Neotropical Fanniidae (Insecta, Diptera): new species of Fannia from Colombia Author Grisales, Diana Author Wolff, Marta Author De, Claudio J. B. text Zootaxa 2012 3591 1 46 journal article 10.5281/zenodo.213946 2a54e127-4d40-404e-ab7f-5b48f6e37e83 1175-5326 213946 Fannia sutagao sp. nov. ( Figs. 19 , 36 , 54 , 72 , 89 , 108 , 126 , 131 ) Diagnosis. these characters apply only to the male sex. Abdomen with a tuft of very long, ventrally developed setae, crossed above the terminalia; hind femur strongly curved, ventral surface with pre-apical conspicuous protuberance with tuft of av and v developed, v with curved apices, 1 long p with curved apex and 2 smaller, basal half setulose; 1 row of pv developed on apical third, not uniform. Description. Holotype male : body length 5.5 mm; wing length 5.5 mm. Head : eye densely setulose. Frontal vitta velvety black. Interocular space ample, 0.22 mm ( Fig. 19 ). Fr 13. Ocellar triangle black; oc proclinate and developed, with 1 pair posterior to posterior ocelli. Poc divergent. Frontoorbital plate and parafacial black, golden pollinose. Parafacial bare, narrowing on ventral third. Fronto-genal suture pronounced and golden-yellowish pollinose. Gena brownish. Lunule black. Scape black, golden pollinose on apical margin, 2 setae reaching pedicel. Pedicel black with dorsal setae developed. Postpedicel black with goldenyellowish pilosity, 2.2 times the length of pedicel. Arista black pubescent. Palpus brown, flattened and claviform. Thorax : black pollinose, weak; scutum without vittae; scutum and scutellum flattened and widened. Acr 2:3 not arranged in rows. Dc 2:3 long. Pprn 2 developed and 1 smaller. Pra 2,longer seta near suture. Prepm setulose. Pre-basal scutellar area setulose, pre-basal seta slightly differentiated from ground setulae. Subapical scutellar setae 1 pair. Discal scutellar setae 1 pair. Lateral scutellar setae present and unordered. Wing : brownish, with upper third between C vein and half of vein R2+3, base of cell br, bm and cup darker. Calypters whitish with margin of upper calypter brown. Haltere brown, with stem base lighter in color. Legs : brown with tarsi dark and pulvilli brown. Fore femur with 1 row of weak av up to median third; 1 row of long pd ; posterior surface densely setulose, with long and strong setae; 1 row of long pv and with straight apices. Ventral surface bare. Fore tibia with 1 pre-apical d ; 1 ad on edge of apical third; anteroventral and ventral surfaces densely setulose from median third to apex, with short setae; 1 pd on edge of apical third and 1 pre-apical; 1 preapical pv; 1 apical v . Fore tarsus with 1 strong v on first tarsomere followed by a thin seta; tarsomeres 3–5 flattened and widened ( Fig. 131 ). Mid femur constricted on pre-apical ventral surface; 1 row of av decreasing in length on basal third where they are more ventral; 1 row of short a ending in longer seta; 1 row of long p up to median third; 1 row of strongly developed pv with hooked apices, ending in 4 longer p ; 1 row of v on basal third, shorter than the pv . Mid tibia on basal half of ventral surface with more pronounced pre-basal constriction, and dorsal surface densely setulose on apical half medially, 1 apical long v , strong and bifid; 1 pre-apical a ; 1 developed on edge of apical third and 2 pre-apical ad , one long and one short, developed; 1 developed pre-apical d ; 1 pd on edge of apical third and 1 pre-apical, strong and long d ; 2 developed pre-apical p . Mid tarsus with 1 short v on first tarsomere. Hind coxa on posterior margin bare, with setae on ventral margin that can be confused with setae on posterior margin. Hind femur strongly curved, on ventral surface with pre-apical conspicuous protuberance, with developed av and v tufts, v with curved apices ( Fig. 36 ); 1 row of long ad on apical half; ventral surface on basal half setulose; 1 long p with curved apex and 2 shorter, basal half setulose; 1 row of pv on apical third, not uniform, and developed ( Fig. 54 ). Hind tibia with 1 median and 1 pre-apical d ; 1 median ad , long and developed; 1 short pre-apical a ; 1 av on edge of apical third and 1 apical; 1 short apical v ; 1 short p apical; 1 apical pv . Hind tarsus missing. Abdomen : elongate, in general with developed setae, brown with syntergite 1+2, tergites 3 and 4 basally translucentbrown, weakly greyish pollinose. Syntergite 1+2 with a set of long lateral setae; tergite 5 with 2 pairs of lateral setae, ventrally with a tuft of long and developed setae on each side that cross above the terminalia. Sternite 1 densely setulose. Sternite 5 as in Fig. 72 . Terminalia ( Figs. 90 , 108 ): epandrium slightly wider than long, with developed setae on basal half; cercal plate elongate apically and bifurcate, with short and weak setae on basal lateral half; surstylus wide, articulated with epandrium, setulose on inner basal margin, apically curved and with short and weak inner setae; bacilliform process short; hypandrium and associated structures as in Fig. 126 . Female : unknown Biology. According to label data, Fannia sutagao sp . nov . is associated with cold ecosystems of the Páramo (up to 3000 m ) and occurs at the Parque Nacional Natural Sumapaz . This ecosystem is characterized by an abundance of water bodies, grasses 40–50 cm long, “frailejón (Asteracea, Espeletia ) and a high diversity of plant and animals (Parques Nacionales Naturales 2002). Comments. these characters apply only to the male sex. Fannia sutagao sp . nov . is morphologically similar to F. iguaque sp . nov . and F. sumapaz sp . nov . , but differs in the strong protuberance on the hind femur, a tuft of developed setae on the protuberance, and a tuft of strong setae on tergite 5 crossing above the terminalia. Etymology. Name in apposition. The species epithet refers to the name the Spaniards gave to the Páramo de Sumapaz , “Páramo de los Sutagaos. This is due to the group of indigenous people who controlled the territory. Type material . Holotype male (IAvH).CO [ Colombia ]. Meta. P.N.N [Parque Nacional Natural] Sumapaz .Bocatoma\ Cerro El Zapato. 4˚14’N 74˚12W\ 3560 m . Malaise 2–17.i.2003 \ A. Patiño \ Leg. M.3442. IAvH. Paratype : CO [ Colombia ]. Meta. P.N.N [Parque Nacional Natural] Sumapaz . Bocatoma\ Cerro El Zapato. 4˚14’N 74˚12W\ 3560 m . Malaise 2–17.i.2003 \ A. Patiño \ Leg. M.3442. IAvH ( 1 male , IAvH). The holotype is a little crushed; the frontal setae and tarsi of both hind legs are missing. Distribution. Colombia : department of Meta (Amazon region).