<document id="B0628B91409E2BAB7C6047E701C98FE2" ID-CLB-Dataset="28153" ID-DOI="10.11646/zootaxa.4563.1.6" ID-GBIF-Dataset="0d849a4d-5dd1-4f60-9036-939f4486fe72" ID-ISSN="1175-5326" ID-Zenodo-Dep="2601017" ID-ZooBank="8E2935BD-3608-4BE1-B565-AE50966EE510" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553171272592" checkinUser="plazi" docAuthor="Martins, Ananda Regina P., Duarte, Marcelo &amp; Robbins, Robert K." docDate="2019" docId="0390D93D7B05FFACFF5BFF17FCE57D5E" docLanguage="en" docName="zootaxa.4563.1.6.pdf" docOrigin="Zootaxa 4563 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Theritas Robbins" docType="treatment" docVersion="7" lastPageNumber="124" masterDocId="FFA9A1457B00FFA9FFCCFF8FFFB87C05" masterDocTitle="Phylogenetic Classification of the Atlides Section of the Eumaeini (Lepidoptera, Lycaenidae)" masterLastPageNumber="134" masterPageNumber="119" pageNumber="124" updateTime="1698703033999" updateUser="ExternalLinkService">
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<mods:title id="51C1FAEB8D73D94D87B6E76AAA752BA7">Phylogenetic Classification of the Atlides Section of the Eumaeini (Lepidoptera, Lycaenidae)</mods:title>
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<mods:namePart id="424426BF70D6776C84949027B52828A9">Martins, Ananda Regina P.</mods:namePart>
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<mods:namePart id="FF98B5A01545148744B54396C8BD219E">Duarte, Marcelo</mods:namePart>
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<mods:title id="0CBCF65FB37847F49BD2DC5131610B4C">Zootaxa</mods:title>
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<mods:date id="DB9DF557377DCDC82D2723080816E900">2019</mods:date>
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<mods:identifier id="1492095924D0C2035B28B0B453B3F105" type="DOI">10.11646/zootaxa.4563.1.6</mods:identifier>
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<treatment id="0390D93D7B05FFACFF5BFF17FCE57D5E" ID-DOI="http://doi.org/10.5281/zenodo.5932754" ID-GBIF-Taxon="154474489" ID-Zenodo-Dep="5932754" LSID="urn:lsid:plazi:treatment:0390D93D7B05FFACFF5BFF17FCE57D5E" httpUri="http://treatment.plazi.org/id/0390D93D7B05FFACFF5BFF17FCE57D5E" lastPageNumber="124" pageId="5" pageNumber="124">
<subSubSection id="C3233BA07B05FFACFF5BFF17FCE57D5E" pageId="5" pageNumber="124" type="nomenclature">
<paragraph id="8B86682B7B05FFACFF5BFF17FD0A7CB7" blockId="5.[151,690,152,178]" box="[151,690,152,178]" pageId="5" pageNumber="124">
<heading id="D0CEDF477B05FFACFF5BFF17FD0A7CB7" bold="true" box="[151,690,152,178]" fontSize="11" level="1" pageId="5" pageNumber="124" reason="1">
<emphasis id="B94DB4397B05FFACFF5BFF17FD0A7CB7" bold="true" box="[151,690,152,178]" pageId="5" pageNumber="124">
<taxonomicName id="4C3913A87B05FFACFF5BFF17FE0E7CB7" ID-CoL="9CM8R" authority="sensu Robbins" authorityName="Robbins" box="[151,438,152,178]" class="Insecta" family="Lycaenidae" genus="Theritas" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="124" phylum="Arthropoda" rank="genus">
<emphasis id="B94DB4397B05FFACFF5BFF17FF447CB7" bold="true" box="[151,252,152,178]" italics="true" pageId="5" pageNumber="124">Theritas</emphasis>
sensu Robbins
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is not monophyletic
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</heading>
</paragraph>
<paragraph id="8B86682B7B05FFACFF5BFF51FCE57D5E" blockId="5.[151,1436,222,347]" pageId="5" pageNumber="124">
<bibRefCitation id="EFA815DA7B05FFACFF5BFF51FEEF7CF2" author="Robbins, R. K." box="[151,343,222,247]" pageId="5" pageNumber="124" pagination="118 - 137" refId="ref9998" refString="Robbins, R. K. (2004 b) Lycaenidae. Theclinae. Tribe Eumaeini. In: Lamas, G. (Ed.), Checklist: Part 4 A. Hesperioidea - Papilionoidea. In: Heppner, J. B. (Ed.), Atlas of Neotropical Lepidoptera. Folume 5 A. Association for Tropical Lepidoptera, Scientific Publishers, Gainesville, pp. 118 - 137." type="book chapter" year="2004">Robbins (2004b)</bibRefCitation>
used 
<taxonomicName id="4C3913A87B05FFACFE52FF51FE437CF2" box="[414,507,222,247]" class="Insecta" family="Lycaenidae" genus="Theritas" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="124" phylum="Arthropoda" rank="genus">
<emphasis id="B94DB4397B05FFACFE52FF51FE437CF2" box="[414,507,222,247]" italics="true" pageId="5" pageNumber="124">Theritas</emphasis>
</taxonomicName>
as a catchall for those 
<taxonomicName id="4C3913A87B05FFACFCC1FF51FCE37CF2" box="[781,859,222,247]" class="Insecta" family="Lycaenidae" genus="Atlides" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="124" phylum="Arthropoda" rank="genus">
<emphasis id="B94DB4397B05FFACFCC1FF51FCE37CF2" box="[781,859,222,247]" italics="true" pageId="5" pageNumber="124">Atlides</emphasis>
</taxonomicName>
Section species that did not belong to the readily recognized 
<taxonomicName id="4C3913A87B05FFACFED6FE8FFEE27D12" authorityName="Swainson" authorityYear="1832" box="[282,346,256,279]" class="Insecta" family="Lycaenidae" genus="Arcas" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="124" phylum="Arthropoda" rank="genus">
<emphasis id="B94DB4397B05FFACFED6FE8FFEE27D12" box="[282,346,256,279]" italics="true" pageId="5" pageNumber="124">Arcas</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C3913A87B05FFACFEA6FF70FDAC7D1D" authorityName="Wallengren" authorityYear="1858" box="[362,532,255,280]" class="Insecta" family="Lycaenidae" genus="Pseudolycaena" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="124" phylum="Arthropoda" rank="genus">
<emphasis id="B94DB4397B05FFACFEA6FF70FDAC7D1D" box="[362,532,255,280]" italics="true" pageId="5" pageNumber="124">Pseudolycaena</emphasis>
</taxonomicName>
, or 
<taxonomicName id="4C3913A87B05FFACFD8EFF70FD287D1D" box="[578,656,255,280]" class="Insecta" family="Lycaenidae" genus="Atlides" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="124" phylum="Arthropoda" rank="genus">
<emphasis id="B94DB4397B05FFACFD8EFF70FD287D1D" box="[578,656,255,280]" italics="true" pageId="5" pageNumber="124">Atlides</emphasis>
</taxonomicName>
. It is polyphyletic in our results (
<figureCitation id="130274AE7B05FFACFBC2FF70FBEC7D1D" box="[1038,1108,255,280]" captionStart="FIGURE 8" captionStartId="4.[151,250,1953,1975]" captionText="FIGURE 8. Cladogram modified from Martins et al. (2018)." pageId="5" pageNumber="124">Fig. 8</figureCitation>
). To maintain the taxonomic stability of 
<taxonomicName id="4C3913A87B05FFACFEEDFEADFED97D3C" authorityName="Swainson" authorityYear="1832" box="[289,353,290,313]" class="Insecta" family="Lycaenidae" genus="Arcas" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="124" phylum="Arthropoda" rank="genus">
<emphasis id="B94DB4397B05FFACFEEDFEADFED97D3C" box="[289,353,290,313]" italics="true" pageId="5" pageNumber="124">Arcas</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C3913A87B05FFACFEB9FEAEFDA67D3F" authorityName="Wallengren" authorityYear="1858" box="[373,542,289,314]" class="Insecta" family="Lycaenidae" genus="Pseudolycaena" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="124" phylum="Arthropoda" rank="genus">
<emphasis id="B94DB4397B05FFACFEB9FEAEFDA67D3F" box="[373,542,289,314]" italics="true" pageId="5" pageNumber="124">Pseudolycaena</emphasis>
</taxonomicName>
, and 
<taxonomicName id="4C3913A87B05FFACFDAAFEAEFD0C7D3F" box="[614,692,289,314]" class="Insecta" family="Lycaenidae" genus="Atlides" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="124" phylum="Arthropoda" rank="genus">
<emphasis id="B94DB4397B05FFACFDAAFEAEFD0C7D3F" box="[614,692,289,314]" italics="true" pageId="5" pageNumber="124">Atlides</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C3913A87B05FFACFD04FEAEFC9D7D3F" box="[712,805,289,314]" class="Insecta" family="Lycaenidae" genus="Theritas" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="124" phylum="Arthropoda" rank="genus">
<emphasis id="B94DB4397B05FFACFD04FEAEFC9D7D3F" box="[712,805,289,314]" italics="true" pageId="5" pageNumber="124">Theritas</emphasis>
</taxonomicName>
is partitioned into 
<taxonomicName id="4C3913A87B05FFACFBC2FEAEFBD37D3F" box="[1038,1131,289,314]" class="Insecta" family="Lycaenidae" genus="Theritas" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="124" phylum="Arthropoda" rank="genus">
<emphasis id="B94DB4397B05FFACFBC2FEAEFBD37D3F" box="[1038,1131,289,314]" italics="true" pageId="5" pageNumber="124">Theritas</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C3913A87B05FFACFBB3FEADFE707D5E" authority="K. Johnson and Lucilda d'Abrera &amp; Balint." authorityName="K. Johnson and Lucilda d'Abrera &amp; Balint." class="Insecta" family="Lycaenidae" genus="Denivia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="124" phylum="Arthropoda" rank="genus">
<emphasis id="B94DB4397B05FFACFBB3FEADFB607D3C" box="[1151,1240,290,313]" italics="true" pageId="5" pageNumber="124">Denivia</emphasis>
K. Johnson and 
<emphasis id="B94DB4397B05FFACFF5BFECDFF557D5E" box="[151,237,322,347]" italics="true" pageId="5" pageNumber="124">Lucilda</emphasis>
d’Abrera &amp; Bálint.
</taxonomicName>
New generic names are not needed.
</paragraph>
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