New Cenozoic Muricidae (Mollusca: Gastropoda) from Europe Author Merle, Didier Author Pacaud, Jean-Michel Author Ledon, Daniel Muséum national d’histoire naturelle, Département Origines et Évolution (CR 2 P - MNHN, CNRS, UPMC, Sorbonne Université), 57 rue Cuvier, 75231 Paris cedex 05 (France) didier. merle @ mnhn. fr (corresponding author) jean-michel. pacaud @ mnhn. fr dnledon @ yahoo. fr didier.merle@mnhn.fr Author Goret, Bernard Raoul Ponchon, F- 31500 Toulouse (France) bernardgoret @ hotmail. com bernardgoret@hotmail.com text Geodiversitas 2024 2024-10-24 46 15 495 551 https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/g2024v46a15.pdf journal article 10.5252/geodiversitas2024v46a15 1638-9395 13985760 urn:lsid:zoobank.org:pub:C32EAAFD-85D8-45CD-8DF7-B894E1537713 Trubatsa calviniacensis n. sp. ( Figs 5G, H ; 25L ) urn:lsid:zoobank.org:act: 287FDBCE-A766-4813-9403-2ABDD9B439E3 Trubatsa parisiensis – Merle & Pacaud 2019: 16 , 17, text-fig 3.5 [ non d’Orbigny, 1850]. TYPE MATERIAL . — Holotype . France • Paris Basin , Oise , Cauvigny ( Châteaurouge ); Calcaire grossier moyen (biozone NP15); middle Eocene (middle Lutetian ); MNHN .F.A25590 ( Pacaud coll.), H: 17 mm ( Figs 5G, H ; 25L ). ETYMOLOGY . — From Calviniacus , the Latin name of the type locality Cauvigny (Oise, France ). TYPE HORIZON . — Calcaire grossier moyen (biozone NP15), see Gély & Lorenz (1991 : pl. 1); middle Eocene (middle Lutetian). TYPE LOCALITY . — France , Paris Basin, Oise, Cauvigny (Châteaurouge), see Fritel 1910: 85 ). DISTRIBUTION . — Only known from the type locality. DESCRIPTION Bulbous protoconch of two whorls; pointed apex.Teleoconch up to 17.8 mm in height, up to 9.2 mm in width, biconic in profile, composed of six whorls. High spire. Last whorl representing up to 70% total height. Spiral angle 62°. Suture weakly impressed. Axial sculpture consisting of four slightly spiny varices per whorl. On spire, anal tube (P1) and short spine (P2). Anal tube forming angle of approximately 45° with shell axis. Anal tube (P1) totally integrated in varices eliminating shoulder. Short P2 spine on first whorls. No callus at base of anal tube. On convex part of last whorl and abapically to anal tube, five spinelets (P2, P3, P4, P5 and P6). Intervarical spaces with weak P2 to P5 cords. Ovate aperture up to 30% of diameter, up to 74% of height of last whorl (including siphonal canal). Edge, erect, smooth. peristome complete. Siphonal canal closed, occupying 47% of apertural lenght. FIG . 6. — Neotype, historical figures, spiral cords and denticles of the outer lip in T. tripteroides ( Lamarck, 1822 ) : A -D , morphotype with rounded varices and poorly distinguishable P1 spine; E -H , morphotype with easily distinguishable P1 spine: A , original figure of Lamarck (1805 : pl. 3, fig. 4); B , C , neotype designated herein, MNHN.F.A90546 (Club géologique d'Île-de-France leg.), Thiverval-Grignon (Falunière), Yvelines, Paris Basin, France, middle Eocene (Lutetian); D , MNHN.F.A90552 (Pacaud coll.), Chaussy (Les Garennes),Paris Basin, France,middle Eocene (Lutetian); E , F , figured specimen of Deshayes (1835 : pl. 82, fig. 1-2); G , H , figured specimen of Cossmann & Pissarro (1911 : pl. 35, fig. 169-1; MNHN.F.J10350 (Cossmann coll.), Chaussy (Les Garennes). Scale bars: 20 mm. Credits: B-D, P. Loubry MNHN/CNRS; G, H, J. Mouchart. COMPARISONS Merle & Pacaud (2019 : pl. 3, fig. 4) attributed a specimen (MNHN.F.A25590) from the middle Lutetian of Cauvigny (Paris Basin) to T. parisiensis (d’Orbigny, 1850) . Like T. ganensis n. sp. it has a paucispiral protoconch and lacks partition. Therefore, it can easily be distinguished from T. parisiensis from the Hampshire Basin which has a multispiral protoconch. Trubatsa calviniacensis n. sp. differs from T. ganensis n. sp. in being higher spired, the primary cord P6 on the last whorl, weak primary cords P2 to P 4 in the intervarical spaces, and by having a more ovate aperture. P2 spine is also more marked on the first teleoconch whorls. Although we have only one specimen , the number of shell differences with the Gan specimens and its occurrence in a strongly different paleoenvironment (deep water at Gan ( Merle 1985 ; Merle & Roux 2018 ) in Aquitaine Basin, versus shallow water in the Paris Basin ( Gély 2008 ) and also quite different age, highly suggest that it represents a different species.