Studies on species of Holarctic Pardosa groups (Araneae, Lycosidae). VIII. The Palearctic species of the Pardosa nigra group Author Kronestedt, Torbjörn Author Marusik, Yuri M. Author Omelko, Mikhail M. Gornotayozhnaya Station, FEB RAS, Gornotayozhnoye, Ussuriysky Dist., Primorsky Kray, 692533, Russia. text Zootaxa 2014 2014-12-11 3894 1 33 60 journal article 269423 http://dx.doi.org/10.11646/zootaxa.3894.1.5 08180ae8-af72-495c-aa70-17a3f17025ec 1175-5326 4956512 0E0A629E-3153-45A7-A929-1DEE66A62252 Pardosa fomichevi sp. n. Figs 2 , 10 , 17–18 , 21 , 29 , 37 , 46–47 , 60, 68, 77–78, 91, 100–101 , 129–130 Type material . Holotype ♂ and allotype ♀ from Russia : Primorsky Kray , Krasnoarmeisk District , slope of Ozernoe Plateau , glades near road, 1359 m ( 45º50’46”N 136º37’31”E ) ( M.M. Omelko ), in ZMMU .— Paratypes . RUSSIA . Primorsky Kray : same data as holotype ( ISEA , NHRS ), 2♂ . Etymology. The specific epithet is a patronym for our colleague Alexander Fomichev from Novosibirsk . Diagnosis. Males can be distinguished from all other members of this group by the curvature of the embolus ( Figs 46–47 ); females by the lateral elevations in the posterior part of the epigyne being at some distance from the septum and their margins diverging at the rear ends ( Fig. 18 ). Description. Male ( holotype ): Total length 6.2. Carapace 3.10 long, 2.35 wide. Prosoma. Carapace brownish. Median band yellowish, narrowed at cephalic-thoracic junction, wide behind PLEs. Lateral bands indistinct, yellowish brown, narrow, broken. Carapace with dark and adpressed light hairs on thoracic sides, median band with adpressed white hairs. Clypeus light brownish. Chelicerae light brownish with longitudinal darker streaks and yellow inside. Sternum dark greyish brown with narrow light stripe in front, furnished with light hairs. Eyes . Width of row I (slightly procurved) 48, row II 69, row III 94, row II-III 67. Diameter of AME 10, ALE 9, PME 25, PLE 21. Distance between AME 5, between AME and ALE 3. Opisthosoma. Dorsum brownish, covered with adpressed light hairs and more erect black hairs. Anterior part and lanceolate stripe with dense light pubescence. Dorsum patterned with black, notably laterally, forming two pairs of black blotches. Venter brown with adpressed light pubescence and numerous scattered short erect hairs. Spinnerets partly sooty with short black hairs. Legs ( Table 1 ). Yellowish. Femora more or less pseudoannulated (=blotches and incomplete transverse markings) in greyish, Fe I and II with proximal half dark greyish. Ti III and IV with very weak darker annulation. Ti I with 3 pairs of ventral spines and two retrolateral spines. TABLE 1. Leg I–IV measurements (mm) of Pardosa fomichevi sp. n. , P. lyrata and P. paramushirensis .

Pardosa fomichevi Male
Fe	Pt	Ti	Mt	Ta	Total
I	2.55	1.10	2.15	2.40	1.55	9.75
II	2.45	1.05	1.95	2.35	1.40	9.20
III	2.30	1.00	1.80	2.55	1.30	8.95
IV	3.00	1.10	2.40	3.90	1.75	12.15
Female
Fe	Pt	Ti	Mt	Ta	Total
I	2.80	1.25	2.30	2.30	1.40	10.05
II	2.70	1.20	2.10	2.30	1.35	9.65
III	2.60	1.10	1.95	2.65	1.30	9.60
IV	3.50	1.30	2.85	4.45	1.80	13.90
Pardosa lyrata Male
Fe	Pt	Ti	Mt	Ta	Total
I	2.50	1.10	2.40	2.65	1.60	10.25
II	2.40	1.05	2.05	2.50	1.40	9.40
III	2.30	1.00	1.80	2.60	1.30	9.00
IV	3.00	1.10	2.50	4.10	1.75	12.45
Female
Fe	Pt	Ti	Mt	Ta	Total
I	2.70	1.20	2.30	2.30	1.35	9.85
II	2.60	1.15	2.05	2.25	1.30	9.35
III	2.45	1.05	1.90	2.60	1.25	9.25
IV	3.25	1.20	2.75	4.20	1.70	13.10
Pardosa paramushirensis Male
Fe	Pt	Ti	Mt	Ta	Total
I	2.50	1.10	2.05	2.30	1.45	9.40
II	2.35	1.05	1.85	2.20	1.35	8.80
III	2.30	1.00	1.70	2.45	1.30	8.75
IV	3.00	1.10	2.35	3.80	1.75	12.00
Female
Fe	Pt	Ti	Mt	Ta	Total
I	2.50	1.25	2.10	2.10	1.35	9.30
II	2.50	1.20	1.85	2.10	1.30	8.95
III	2.40	1.10	1.85	2.40	1.25	9.00
IV	3.15	1.25	2.70	3.95	1.70	12.75

FIGURES 20–27. Bulbus of left male palp in ventral view. 20, Pardosa eiseni . 21, P. fomichevi sp. n. 22, P. giebeli . 23, P. lasciva . 24, P. lyrata . 25, P. nigra . 26, P. paramushirensis . 27, P. trailli . Scale lines 0.2 mm. Palp ( Figs 2 , 10 , 17 , 21 , 29 ). Pt 0.65, Ti 0.60, Cy 1.35. Brownish with darker markings. Longitudinal darker markings on Ti with with numerous strong black hairs. Cy blackish, distally light brown to yellowish. Tegular apophysis with ¾ of prolaterad rim of anteriorly directed branch very slightly concave ( Figs 17 , 22 , 37 ). Conductor as in Fig 78 , terminal apophysis as in Figs 60 , 68 & 77–78 . Embolus ( Figs 46–47 , 60, 68) wide, screw-shaped, apically curved inwards, narrowed, and tapering to pointed tip. FIGURES 28–35. Bulbus of left male palp in oblique ventral-frontal view. 28, Pardosa eiseni . 29, P. fomichevi sp. n. 30, P. giebeli . 31, P. lasciva . 32, P. lyrata . 33, P. nigra . 34, P. paramushirensis . 35, P. trailli . Scale lines 0.2 mm. Female ( allotype ). Total length 6.8. Carapace 3.40 long, 2.60 wide. Prosoma and opisthosoma . Carapace similar to that in the male, but with lateral bands more distinct. Opisthosoma patterned as in the male. Venter of opisthosoma brown with adpressed white pubescence. Eyes. Width of row I (slightly procurved) 54, row II 76, row III 104, row II-III 74. Diameter of AME 11, ALE 10, PME 26, PLE 22. Distance between AME 7, between AME and ALE 4. Legs ( Table 1 ). Yellowish, clearly annulated in grey (femora) and light brown (patellae-metatarsi). Ti I with two retrolateral spines. Epigyne ( Figs 18 , 91 ). Lateral elevations diverging backwards (45º) from about half the length of the epigyne. Septum only partly filling the cavities in the posterior half. Spermathecae ovoid ( Figs 100–101 ). Size variation. Carapace length: males 2.95–3.10 (n=3). Distribution ( Figs 129–130 ). Russia : Primorsky Kray (only known from the type locality).