Systematics of Opsiphanes Doubleday, [1849] (Lepidoptera: Nymphalidae, Satyrinae, Brassolini): an integrative approach Author Piovesan, Mônica 0000-0003-3367-9358 monica367piovesan@gmail.com Author Casagrande, Mirna Martins 0000-0002-6076-8463 mibras@ufpr.br Author Mielke, Olaf Hermann Hendrik 0000-0003-3655-4606 omhesp@ufpr.br text Zootaxa 2022 2022-12-06 5216 1 1 278 http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN journal article 10.11646/zootaxa.5216.1.1 1175-5326 7403532 F9C92B91-42CF-4D2D-B315-14FF801815D4 Opsiphanes fabricii ( Boisduval, 1870 ) ( Figs 169–186 ) Diagnosis. The male of Opsiphanes fabricii differs from the males of Opsiphanes jacobsorum and Opsiphanes periphetes stat. nov. in the DHW with a defined submarginal band extending to the anal region, usually as wide as the DFW oblique band, between CuA 1 and 2A. The female of Opsiphanes fabricii , on the other hand, differs from the female of Opsiphanes jacobsorum in the yellow DHW submarginal band, and from the female of Opsiphanes periphetes stat. nov. in the continuous DHW submarginal band between the veins, from the costal margin to, usually, M 2 ; the DHW submarginal band has the same width as the DFW oblique band ( Figs 169a , 171a ). Opsiphanes fabricii and Opsiphanes merianae stat. rest. are practically indistinguishable by the external morphology, and are differentiated by means of the genitalia. While the male of Opsiphanes fabricii has the tegumen in dorsal view with an anteriorly projected margin, in the male of Opsiphanes merianae stat. rest. the anterior margin of the tegumen has a slight median concavity. The female of Opsiphanes fabricii has a C-shaped signum while Opsiphanes merianae stat. rest. has two parallel signa. The geographical distribution can also be used to separate the two species; while Opsiphanes fabricii subspecies are trans-Andean, Opsiphanes merianae stat. rest. subspecies are cis-Andean. Opsiphanes fabricii is closely related to Opsiphanes jacobsorum , Opsiphanes merianae stat. rest. and Opsiphanes periphetes stat. nov. (genetic distance 0.9–1.2%, 1.2–1.9% and 1.0–1.7%, respectively). Male genitalia. Tegumen in dorsal view with anterior margin anteriorly projected, shorter than the uncus; anterior projection of saccus in lateral view twice as long as dorsal arms of saccus ( Fig. 179 ). Female genitalia. Lamella antevaginalis membranous medially and sclerotized laterally; lamella postvaginalis rounded medially; ostium with weaker sclerotization than lamella ante and postvaginalis; corpus bursae longer than the ductus bursae, with C-shaped signum ( Fig. 180 ). Variation. Subspecies vary in the following characters: DFW, the width of the band inside the discal cell, and in the recesses of the anterior and posterior margins of this band; the width and length of the oblique band; due to variations in the width of these bands, the bifurcated appearence is not always continuous. DHW, the width of the submarginal band. VHW, the zigzag submarginal line EIII is not always evident ( Figs 169–178 ).