<document id="0C90F9BD6CBECF5422BAD679B36F5DB8" ID-DOI="10.11646/zootaxa.5383.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="10301516" ID-ZooBank="7A1B3C63-D41A-4F5D-976D-BEE4B8B02ABE" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1702024299473" checkinUser="plazi" docAuthor="Oliveira, Luiz Fernando M., De Alvarenga, José Geraldo G. &amp; Brescovit, Antonio D." docDate="2023" docId="CB4F87E5996FFFFA278D928B33D9F85A" docLanguage="en" docName="zootaxa.5383.1.1.pdf" docOrigin="Zootaxa 5383 (1)" docSource="https://www.mapress.com/zt/article/download/zootaxa.5383.1.1/52436" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Otoniela adisi Brescovit 1997" docType="treatment" docVersion="2" lastPageNumber="3" masterDocId="3776FF9D996DFFF8271A94693328FFA3" masterDocTitle="On the Neotropical spider genus Otoniela Brescovit, 1997 (Araneae: Anyphaenidae, Anyphaeninae), with the description of six new species" masterLastPageNumber="23" masterPageNumber="1" pageNumber="3" updateTime="1702036840052" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="E4D9F9D5FC92431D67E4928DB77CDF4C">On the Neotropical spider genus Otoniela Brescovit, 1997 (Araneae: Anyphaenidae, Anyphaeninae), with the description of six new species</mods:title>
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<mods:namePart id="FD49C637E18D74721DF4CF93FDD87E1A">Oliveira, Luiz Fernando M.</mods:namePart>
<mods:affiliation id="12FDCB4D304550711596AE81D97C30D2">Laboratório de Coleções Zoológicas, Instituto Butantan, Av. Vital Brasil, 1500, 05503 - 900, São Paulo, SP, Brazil</mods:affiliation>
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<mods:namePart id="05C3BF93EA51E225955A004E6C9D8151">De Alvarenga, José Geraldo G.</mods:namePart>
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<mods:affiliation id="F0C58363ED95C396476FE3AD98FBEE57">Laboratório de Coleções Zoológicas, Instituto Butantan, Av. Vital Brasil, 1500, 05503 - 900, São Paulo, SP, Brazil &amp; ge. alvarenga. bio @ gmail. com; https: // orcid. org / 0000 - 0003 - 2864 - 6981</mods:affiliation>
<mods:nameIdentifier id="60EB32C80B498F8FFAA833CB23F53FC4" type="email">ge.alvarenga.bio@gmail.com</mods:nameIdentifier>
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<mods:namePart id="6BE3E5F4429A973C8E76E618EE646025">Brescovit, Antonio D.</mods:namePart>
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<mods:affiliation id="6156530F72F40B22737D83CDE94AC49E">Laboratório de Coleções Zoológicas, Instituto Butantan, Av. Vital Brasil, 1500, 05503 - 900, São Paulo, SP, Brazil &amp; antonio. brescovit @ butantan. gov. br; https: // orcid. org / 0000 - 0002 - 1511 - 5324</mods:affiliation>
<mods:nameIdentifier id="180559803E782AAD3FE735BE7EF91A37" type="email">antonio.brescovit@butantan.gov.br</mods:nameIdentifier>
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<treatment id="CB4F87E5996FFFFA278D928B33D9F85A" LSID="urn:lsid:plazi:treatment:CB4F87E5996FFFFA278D928B33D9F85A" httpUri="http://treatment.plazi.org/id/CB4F87E5996FFFFA278D928B33D9F85A" lastPageNumber="3" pageId="2" pageNumber="3">
<subSubSection id="0BFC6578996FFFFA278D928B3202F95E" box="[151,298,1762,1789]" pageId="2" pageNumber="3" type="nomenclature">
<paragraph id="435936F3996FFFFA278D928B3202F95E" blockId="2.[151,298,1762,1789]" box="[151,298,1762,1789]" pageId="2" pageNumber="3">
<heading id="1811819F996FFFFA278D928B3202F95E" bold="true" box="[151,298,1762,1789]" fontSize="11" level="1" pageId="2" pageNumber="3" reason="1">
<emphasis id="7192EAE1996FFFFA278D928B3202F95E" bold="true" box="[151,298,1762,1789]" pageId="2" pageNumber="3">
Group 
<taxonomicName id="84E64D70996FFFFA27EB928B3202F95E" ID-CoL="4B7LH" authorityName="Brescovit" authorityYear="1997" box="[241,298,1762,1789]" class="Arachnida" family="Anyphaenidae" genus="Otoniela" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="adisi">
<emphasis id="7192EAE1996FFFFA27EB928B3202F95E" bold="true" box="[241,298,1762,1789]" italics="true" pageId="2" pageNumber="3">adisi</emphasis>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="0BFC6578996FFFFA278D934333D9F85A" pageId="2" pageNumber="3" type="diagnosis">
<paragraph id="435936F3996FFFFA278D9343372FF812" blockId="2.[151,1437,1834,2041]" pageId="2" pageNumber="3">
<emphasis id="7192EAE1996FFFFA278D93433222F8E7" bold="true" box="[151,266,1834,1860]" pageId="2" pageNumber="3">Diagnosis</emphasis>
. The 
<taxonomicName id="84E64D70996FFFFA2650934232A8F8E6" authorityName="Brescovit" authorityYear="1997" box="[330,384,1835,1861]" class="Arachnida" family="Anyphaenidae" genus="Otoniela" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="adisi">
<emphasis id="7192EAE1996FFFFA2650934232A8F8E6" box="[330,384,1835,1861]" italics="true" pageId="2" pageNumber="3">adisi</emphasis>
</taxonomicName>
group differs from the 
<taxonomicName id="84E64D70996FFFFA259E9342303EF8E6" baseAuthorityName="Simon" baseAuthorityYear="1896" box="[644,790,1835,1861]" class="Arachnida" family="Anyphaenidae" genus="Otoniela" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="quadrivittata">
<emphasis id="7192EAE1996FFFFA259E9342303EF8E6" box="[644,790,1835,1861]" italics="true" pageId="2" pageNumber="3">quadrivittata</emphasis>
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group by the palp with a absence of prolaterally modified cymbial spine, transversal triangular ventral tegular projection with wide base and pointed apex, embolus short, with a basal embolar apophysis; female epigynum with lateral borders of anterior field inconspicuous and internally with short copulatory ducts (
<figureCitation id="DBDD2A76996FFFFA26C293FE316FF812" box="[472,583,1943,1969]" captionStart="FIGURE 1" captionStartId="3.[151,250,1690,1714]" captionTargetBox="[289,1298,179,1664]" captionTargetId="figure-19@3.[289,1298,179,1664]" captionTargetPageId="3" captionText="FIGURE 1A–J. Otoniela adisi Brescovit, 1997.Male (IBSP 29715; 58258):A habitus, dorsal view; C Cephalothorax, chelicerae, dorsal view (arrow indicate fang base with protuberance); D left palp expanded, prolateral view; E left palp expanded, retrolateral view; F left palp, ventral view; G left palp, prolateral view; H left palp, retrolateral view; I left palp, retrodorsal view. Female (IBSP 11452): B habitus, dorsal view; J Epigynum, ventral view. Abbreviations:A, atrium; EBA, embolar base apophysis; BH, basal haematodocha; CO, copulatory opening; C, cymbium; DPA, dorsal patellar apophysis; E, embolus; LB, lateral borders; MA, median apophysis; P, petiole; RTA, retrolateral tibial apophysis; PTA, prolateral tibial apophysis; RPA, retrolateral patellar apophysis; ST, subtegulum; T, tegulum; VTP, ventral tegular projection. Scale bars: A, B 2.16mm; C 0.83mm; D, E, F, J 0.25mm; G–I 0.5mm." figureDoi="http://doi.org/10.5281/zenodo.10301518" httpUri="https://zenodo.org/record/10301518/files/figure.png" pageId="2" pageNumber="3">Figs 1F, J</figureCitation>
; 
<figureCitation id="DBDD2A76996FFFFA254993FE31B8F812" box="[595,656,1943,1969]" captionStart="FIGURE 3" captionStartId="5.[151,250,1682,1706]" captionTargetBox="[292,1296,181,1658]" captionTargetId="figure-19@5.[292,1296,181,1658]" captionTargetPageId="5" captionText="FIGURE 3A–L. Otoniela adisi Brescovit, 1997. Male (IBSP 11452):A left palp, ventral view; B same, prolateral view; C same, ventral view; D same, retrolateral view (arrows indicate lyriform organ in the patella); E left palp, patella, retrolateral view (arrows indicate the lyriform organ in the patella); F left palp, dorsal view; G left palp, copulatory bulb, ventral view (arrow indicate the denticle in the median apophysis); H same, ventral view; I same, retroventral view (arrow indicate the denticle in the median apophysis). Female (IBSP 11454): J vulva, dorsal view; K same, zoom in spermatheca, dorsal view; L vulva, details spermatheca, dorsal view. Abbreviations: EBA, embolar base apophysis; BG, Bennett’s gland, CD, copulatory ducts; DPA, dorsal patellar apophysis; E, embolus; FD, fertilization ducts; GD, gland ducts; MA, median apophysis; PTA, prolateral tibial apophysis; RTA, retrolateral tibial apophysis; RPA, retrolateral patellar apophysis; S, spermathecae; SR, seminal receptacles; ST, subtegulum; T, tegulum; VTP, ventral tegular projection. Scale bars: A 0.4mm; B, G, J 0.2mm; C, D, F, H, I, K 0.01mm; E 0.05mm; L 0.03mm." figureDoi="http://doi.org/10.5281/zenodo.10301528" httpUri="https://zenodo.org/record/10301528/files/figure.png" pageId="2" pageNumber="3">3G–I</figureCitation>
; 
<figureCitation id="DBDD2A76996FFFFA258693FE31F5F812" box="[668,733,1943,1969]" captionStart="FIGURE 4" captionStartId="7.[151,250,1832,1856]" captionTargetBox="[191,1395,489,1809]" captionTargetId="figure-153@7.[191,1395,489,1809]" captionTargetPageId="7" captionText="FIGURE 4A–H. Otoniela lupercioi sp. nov.. Male (IBSP 29716): A habitus, dorsal view; C, D left palp, ventral view; E same, prolateral view; F same, retrolateral view. Female (IBSP 29715): B habitus, dorsal view; G epigynum, ventral view; H vulva, dorsal view. Abbreviations: A, atrium; EBA, embolar basal apophysis; CD, copulatory ducts; DPA, dorsal patellar apophysis; E, embolus; FD, fertilization ducts; LB, lateral borders; MA, median apophysis; PTA, prolateral tibial apophysis; RTA, retrolateral tibial apophysis; RPA, retrolateral patellar apophysis; S, spermatheca; ST, subtegulum, T, tegulum, VTP, ventral tegular projection. Scale bars: A 2.6mm; B 3.5mm; C, F, G 0.25mm; D, E 0.5mm." figureDoi="http://doi.org/10.5281/zenodo.10301530" httpUri="https://zenodo.org/record/10301530/files/figure.png" pageId="2" pageNumber="3">4C, F</figureCitation>
; 
<figureCitation id="DBDD2A76996FFFFA25F393FF3018F812" box="[745,816,1942,1969]" captionStart="FIGURE 5" captionStartId="8.[151,250,1752,1776]" captionTargetBox="[268,1319,181,1728]" captionTargetId="figure-24@8.[268,1319,181,1728]" captionTargetPageId="8" captionText="FIGURE 5A–J. Otoniela lupercioi sp. nov.. Male (IBSP 79971): A chelicera, dorsal view; B chelicera, ventral view; C right chelicera, ventral view; D left palp, prolateral view; E same, ventral view; F same retrolateral view (arrow indicate the lyriform organ in the patella); G left palp, copulatory bulb, ventral view; H same, retrolateral view. Female (IBSP 279589): I vulva, dorsal view; J vulva, detail, dorsal view. Abbreviations: EBA, embolar base apophysis; BG, Bennett’s gland, CD, copulatory ducts; DPA, dorsal patellar apophysis; E, embolus; FD, fertilization ducts; GD, gland ducts; MA, median apophysis; PTA, prolateral tibial apophysis; RTA, retrolateral tibial apophysis; RPA, retrolateral patellar apophysis; S, spermathecae; SR, seminal receptacles; ST, subtegulum; T, tegulum; VTP, ventral tegular projection. Scale bars: A, B, D 0.5mm; C, H, I 0.1mm; E, F 0.2mm; G 0.3mm; J 0.04mm." figureDoi="http://doi.org/10.5281/zenodo.10301532" httpUri="https://zenodo.org/record/10301532/files/figure.png" pageId="2" pageNumber="3">5G, H</figureCitation>
; 
<figureCitation id="DBDD2A76996FFFFA242693FE3056F812" box="[828,894,1943,1969]" captionStart="FIGURE 6" captionStartId="10.[151,250,1828,1852]" captionTargetBox="[177,1411,453,1805]" captionTargetId="figure-143@10.[177,1411,453,1805]" captionTargetPageId="10" captionText="FIGURE 6A–H. Otoniela chica sp. nov.. Male (MACN-Ar): A habitus, dorsal view; C, D left palp, ventral view; E same, prolateral view; F same, retrolateral view. Female (MLP): B habitus, dorsal view; G epigynum, ventral view; H vulva, dorsal view.Abbreviations:A, atrium; EBA, embolar base apophysis; CD, copulatory ducts; DPA, dorsal patellar apophysis; E, embolus; FD, fertilization ducts; LB, lateral borders; MA, median apophysis; PTA, prolateral tibial apophysis; RTA, retrolateral tibial apophysis; RPA, retrolateral patellar apophysis; S, spermatheca; ST, subtegulum, T, tegulum; VTP, ventral tegular projection. Scale bars: A, B 2.16mm; C, F, G 0.25mm; D, E 0.6mm." figureDoi="http://doi.org/10.5281/zenodo.10301536" httpUri="https://zenodo.org/record/10301536/files/figure.png" pageId="2" pageNumber="3">6C, F</figureCitation>
; 
<figureCitation id="DBDD2A76996FFFFA249393FE30E3F812" box="[905,971,1943,1969]" captionStart="FIGURE 7" captionStartId="12.[151,250,1756,1780]" captionTargetBox="[177,1411,381,1733]" captionTargetId="figure-115@12.[177,1411,381,1733]" captionTargetPageId="12" captionText="FIGURE 7A–H. Otoniela duovizinhense sp. nov.. Male (IBSP 21060): A habitus, dorsal view; C, D left palp, ventral view; E same, prolateral view; F same, retrolateral view. Female (IBSP 21060): B habitus, dorsal view; G epigynum, ventral view; H vulva, dorsal view. Abbreviations: A, atrium; EBA, embolar base apophysis; CD, copulatory ducts; CO, copulatory opening; DPA, dorsal patellar apophysis; E, embolus; LB, lateral borders; MA, median apophysis; PTA, prolateral tibial apophysis; RTA, retrolateral tibial apophysis; RPA, retrolateral patellar apophysis; S, spermatheca; ST, subtegulum, T, tegulum, VTP, ventral tibial projection. Scale bars: A 2.16mm; B 1.5mm; C 0.25mm; D, E 0.5mm; F, G 0.25mm." figureDoi="http://doi.org/10.5281/zenodo.10301540" httpUri="https://zenodo.org/record/10301540/files/figure.png" pageId="2" pageNumber="3">7C, F</figureCitation>
; 
<figureCitation id="DBDD2A76996FFFFA24CC93FE30D2F812" box="[982,1018,1943,1969]" captionStart="FIGURE 8" captionStartId="13.[151,250,1853,1877]" captionTargetBox="[151,1436,885,1829]" captionTargetId="figure-225@13.[151,1436,885,1830]" captionTargetPageId="13" captionText="FIGURE 8A–D. Otoniela aparecida sp. nov.. Female (IBSP 21105):A habitus, dorsal view; B same, ventral view; C epigynum, ventral view; D vulva, dorsal view. Abbreviations: A, atrium; CD, copulatory ducts; FD, fertilization ducts; LB, lateral borders; S, spermatheca. Scale bars: A 1.5mm; B 2.16mm, C, D 0.25mm." figureDoi="http://doi.org/10.5281/zenodo.10301542" httpUri="https://zenodo.org/record/10301542/files/figure.png" pageId="2" pageNumber="3">8C</figureCitation>
).
</paragraph>
<paragraph id="435936F3996FFFFA27DD93D333D9F85A" blockId="2.[151,1437,1834,2041]" pageId="2" pageNumber="3">
<emphasis id="7192EAE1996FFFFA27DD93D33277F877" bold="true" box="[199,351,1978,2004]" pageId="2" pageNumber="3">Composition</emphasis>
. 
<taxonomicName id="84E64D70996FFFFA267493D232EFF876" authorityName="Brescovit" authorityYear="1997" box="[366,455,1979,2005]" class="Arachnida" family="Anyphaenidae" genus="Otoniela" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="adisi">
<emphasis id="7192EAE1996FFFFA267493D232EFF876" box="[366,455,1979,2005]" italics="true" pageId="2" pageNumber="3">O. adisi</emphasis>
</taxonomicName>
, 
<taxonomicName id="84E64D70996FFFFA26CC93D23175F876" authority="Oliveira &amp; De Alvarenga &amp; Brescovit, 2023" authorityName="Oliveira &amp; De Alvarenga &amp; Brescovit" authorityYear="2023" box="[470,605,1979,2005]" class="Arachnida" family="Anyphaenidae" genus="Otoniela" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="lupercioi" status="sp. nov.">
<emphasis id="7192EAE1996FFFFA26CC93D23175F876" box="[470,605,1979,2005]" italics="true" pageId="2" pageNumber="3">O. lupercioi</emphasis>
</taxonomicName>
<emphasis id="7192EAE1996FFFFA257F93D23197F876" bold="true" box="[613,703,1979,2005]" pageId="2" pageNumber="3">
<taxonomicNameLabel id="6AA1579A996FFFFA257F93D23197F876" box="[613,703,1979,2005]" pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, 
<taxonomicName id="84E64D70996FFFFA25CC93D2301EF876" authority="Oliveira &amp; De Alvarenga &amp; Brescovit, 2023" authorityName="Oliveira &amp; De Alvarenga &amp; Brescovit" authorityYear="2023" box="[726,822,1979,2005]" class="Arachnida" family="Anyphaenidae" genus="Otoniela" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="chica" status="sp. nov.">
<emphasis id="7192EAE1996FFFFA25CC93D2301EF876" box="[726,822,1979,2005]" italics="true" pageId="2" pageNumber="3">O. chica</emphasis>
</taxonomicName>
<emphasis id="7192EAE1996FFFFA242793D230BFF876" bold="true" box="[829,919,1979,2005]" pageId="2" pageNumber="3">
<taxonomicNameLabel id="6AA1579A996FFFFA242793D230BFF876" box="[829,919,1979,2005]" pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, 
<taxonomicName id="84E64D70996FFFFA24B493D23747F876" authority="Oliveira &amp; De Alvarenga &amp; Brescovit, 2023" authorityName="Oliveira &amp; De Alvarenga &amp; Brescovit" authorityYear="2023" box="[942,1135,1979,2005]" class="Arachnida" family="Anyphaenidae" genus="Otoniela" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="duovizinhense" status="sp. nov.">
<emphasis id="7192EAE1996FFFFA24B493D23747F876" box="[942,1135,1979,2005]" italics="true" pageId="2" pageNumber="3">O. duovizinhense</emphasis>
</taxonomicName>
<emphasis id="7192EAE1996FFFFA236D93D237F9F876" bold="true" box="[1143,1233,1979,2005]" pageId="2" pageNumber="3">
<taxonomicNameLabel id="6AA1579A996FFFFA236D93D237F9F876" box="[1143,1233,1979,2005]" pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and 
<taxonomicName id="84E64D70996FFFFA221393D236B5F876" authority="Oliveira &amp; De Alvarenga &amp; Brescovit, 2023" authorityName="Oliveira &amp; De Alvarenga &amp; Brescovit" authorityYear="2023" box="[1289,1437,1979,2005]" class="Arachnida" family="Anyphaenidae" genus="Otoniela" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="aparecida" status="sp. nov.">
<emphasis id="7192EAE1996FFFFA221393D236B5F876" box="[1289,1437,1979,2005]" italics="true" pageId="2" pageNumber="3">O. aparecida</emphasis>
</taxonomicName>
<emphasis id="7192EAE1996FFFFA278D93B633D9F85A" bold="true" box="[151,241,2015,2041]" pageId="2" pageNumber="3">
<taxonomicNameLabel id="6AA1579A996FFFFA278D93B633D9F85A" box="[151,241,2015,2041]" pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>