Revision of Telothyria van der Wulp (Diptera: Tachinidae) and twenty-five new species from Area de Conservacion Guanacaste in northwestern Costa Rica with a key to Mesoamerican species Author Fleming, AJ Agriculture and Agri-Food Canada, Ottawa, Canada https://orcid.org/0000-0002-0943-8047 ajfleming604@gmail.com Author Wood, D. Monty Agriculture and Agri-Food Canada, Ottawa, Canada Author Smith, M. Alex University of Guelph, Guelph, Canada https://orcid.org/0000-0002-8650-2575 Author Dapkey, Tanya University of Pennsylvania, Philadelphia, United States of America Author Hallwachs, Winnie University of Pennsylvania, Philadelphia, United States of America Author Janzen, Daniel University of Pennsylvania, Philadelphia, United States of America text Biodiversity Data Journal 2020 8 47157 47157 http://dx.doi.org/10.3897/BDJ.8.e47157 journal article http://dx.doi.org/10.3897/BDJ.8.e47157 1314-2828-8-e47157 1EB383EF49254AC6821DE622BD60C371 0E84784338D1531C908DFC75AF1E9FC3 Telothyria manuelpereirai Fleming & Wood sp. n. Materials Type status: Holotype . Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0011567 ; recordedBy: D.H. Janzen, W. Hallwachs & Minor Carmona ; individualID: DHJPAR0011567; individualCount: 1 ; sex: M ; lifeStage: adult ; preparations: pinned; otherCatalogNumbers: ASTAQ954-06, 05-SRNP-42650, BOLD:AAB8712; Taxon: scientificName: Telothyriamanuelpereirai; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Telothyria; specificEpithet: manuelpereirai; scientificNameAuthorship: Fleming & Wood, 2018; Location: continent: Central America; country: Costa Rica ; countryCode: CR; stateProvince: Alajuela; county: Sector Rincon Rain Forest; locality: Area de Conservacion Guanacaste ; verbatimLocality: Sendero Tucan; verbatimElevation: 410 ; verbatimLatitude: 10.9042; verbatimLongitude: -85.2712; verbatimCoordinateSystem: Decimal; decimalLatitude: 10.9042 ; decimalLongitude: -85.2712 ; Identification: identifiedBy: AJ Fleming ; dateIdentified: 2018; Event: samplingProtocol: Reared from the larve of the Crambidae, Piletosoma thialisDHJ03 ; verbatimEventDate: 15-Oct-2005 ; Record Level: language: en; institutionCode: CNC ; collectionCode: Insects ; basisOfRecord: Pinned Specimen Type status: Paratype . Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0006599 ; recordedBy: D.H. Janzen, W. Hallwachs & Jose Perez ; individualID: DHJPAR0006599; individualCount: 1 ; sex: F ; lifeStage: adult ; preparations: pinned; otherCatalogNumbers: ASTA778-06, 05-SRNP-43026, BOLD:AAB8712; Taxon: scientificName: Telothyriamanuelpereirai; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Telothyria; specificEpithet: manuelpereirai; scientificNameAuthorship: Fleming & Wood, 2018; Location: continent: Central America; country: Costa Rica ; countryCode: CR; stateProvince: Alajuela; county: Sector Rincon Rain Forest; locality: Area de Conservacion Guanacaste ; verbatimLocality: Sendero Rincon; verbatimElevation: 430 ; verbatimLatitude: 10.8962; verbatimLongitude: -85.2777; verbatimCoordinateSystem: Decimal; decimalLatitude: 10.8962 ; decimalLongitude: -85.2777 ; Identification: identifiedBy: AJ Fleming ; dateIdentified: 2018; Event: samplingProtocol: Reared from the larve of the Crambidae, Piletosoma thialisDHJ03 ; verbatimEventDate: 23-Nov-2005 ; Record Level: language: en; institutionCode: CNC ; collectionCode: Insects ; basisOfRecord: Pinned Specimen Type status: Paratype . Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0006648 ; recordedBy: D.H. Janzen, W. Hallwachs & Minor Carmona ; individualID: DHJPAR0006648; individualCount: 1 ; sex: M ; lifeStage: adult ; preparations: pinned; otherCatalogNumbers: ASTA826-06, 05-SRNP-43897, BOLD:AAB8712; Taxon: scientificName: Telothyriamanuelpereirai; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Telothyria; specificEpithet: manuelpereirai; scientificNameAuthorship: Fleming & Wood, 2018; Location: continent: Central America; country: Costa Rica ; countryCode: CR; stateProvince: Alajuela; county: Sector Rincon Rain Forest; locality: Area de Conservacion Guanacaste ; verbatimLocality: Estacion Caribe; verbatimElevation: 415 ; verbatimLatitude: 10.9019; verbatimLongitude: -85.2749; verbatimCoordinateSystem: Decimal; decimalLatitude: 10.9019 ; decimalLongitude: -85.2749 ; Identification: identifiedBy: AJ Fleming ; dateIdentified: 2018; Event: samplingProtocol: Reared from the larve of the Crambidae, Piletosoma thialisDHJ03 ; verbatimEventDate: 27-Jan-2006 ; Record Level: language: en; institutionCode: CNC ; collectionCode: Insects ; basisOfRecord: Pinned Specimen Type status: Paratype . Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0007127 ; recordedBy: D.H. Janzen, W. Hallwachs & Minor Carmona ; individualID: DHJPAR0007127; individualCount: 1 ; sex: M ; lifeStage: adult ; preparations: pinned; otherCatalogNumbers: ASTAV369-06, 06-SRNP-40650, BOLD:AAB8712; Taxon: scientificName: Telothyriamanuelpereirai; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Telothyria; specificEpithet: manuelpereirai; scientificNameAuthorship: Fleming & Wood, 2018; Location: continent: Central America; country: Costa Rica ; countryCode: CR; stateProvince: Alajuela; county: Sector Rincon Rain Forest; locality: Area de Conservacion Guanacaste ; verbatimLocality: Sendero Tucan; verbatimElevation: 410 ; verbatimLatitude: 10.9042; verbatimLongitude: -85.2712; verbatimCoordinateSystem: Decimal; decimalLatitude: 10.9042 ; decimalLongitude: -85.2712 ; Identification: identifiedBy: AJ Fleming ; dateIdentified: 2018; Event: samplingProtocol: Reared from the larve of the Crambidae, Piletosoma thialisDHJ03 ; verbatimEventDate: 24-Mar-2006 ; Record Level: language: en; institutionCode: CNC ; collectionCode: Insects ; basisOfRecord: Pinned Specimen Type status: Paratype . Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0014977 ; recordedBy: D.H. Janzen, W. Hallwachs & Jose Perez ; individualID: DHJPAR0014977; individualCount: 1 ; sex: M ; lifeStage: adult ; preparations: pinned; otherCatalogNumbers: ASTAV668-06, 06-SRNP-41922, BOLD:AAB8712; Taxon: scientificName: Telothyriamanuelpereirai; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Telothyria; specificEpithet: manuelpereirai; scientificNameAuthorship: Fleming & Wood, 2018; Location: continent: Central America; country: Costa Rica ; countryCode: CR; stateProvince: Alajuela; county: Sector Rincon Rain Forest; locality: Area de Conservacion Guanacaste ; verbatimLocality: Sendero Tucan; verbatimElevation: 410 ; verbatimLatitude: 10.9042; verbatimLongitude: -85.2712; verbatimCoordinateSystem: Decimal; decimalLatitude: 10.9042 ; decimalLongitude: -85.2712 ; Identification: identifiedBy: AJ Fleming ; dateIdentified: 2018; Event: samplingProtocol: Reared from the larve of the Crambidae, Piletosoma thialisDHJ02 ; verbatimEventDate: 29-Jun-2006 ; Record Level: language: en; institutionCode: CNC ; collectionCode: Insects ; basisOfRecord: Pinned Specimen Type status: Paratype . Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0014987 ; recordedBy: D.H. Janzen, W. Hallwachs & Minor Carmona ; individualID: DHJPAR0014987; individualCount: 1 ; sex: F ; lifeStage: adult ; preparations: pinned; otherCatalogNumbers: ASTAV678-06, 06-SRNP-42052, BOLD:AAB8712; Taxon: scientificName: Telothyriamanuelpereirai; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Telothyria; specificEpithet: manuelpereirai; scientificNameAuthorship: Fleming & Wood, 2018; Location: continent: Central America; country: Costa Rica ; countryCode: CR; stateProvince: Alajuela; county: Sector Rincon Rain Forest; locality: Area de Conservacion Guanacaste ; verbatimLocality: Sendero Anonas; verbatimElevation: 405 ; verbatimLatitude: 10.9053; verbatimLongitude: -85.2788; verbatimCoordinateSystem: Decimal; decimalLatitude: 10.9053 ; decimalLongitude: -85.2788 ; Identification: identifiedBy: AJ Fleming ; dateIdentified: 2018; Event: samplingProtocol: Reared from the larve of the Crambidae, Piletosoma thialisDHJ02 ; verbatimEventDate: 06-Jul-2006 ; Record Level: language: en; institutionCode: CNC ; collectionCode: Insects ; basisOfRecord: Pinned Specimen Type status: Paratype . Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0010354 ; recordedBy: D.H. Janzen, W. Hallwachs & Minor Carmona ; individualID: DHJPAR0010354; individualCount: 1 ; sex: F ; lifeStage: adult ; preparations: pinned; otherCatalogNumbers: ASTAS185-06, 06-SRNP-42553, BOLD:AAB8712; Taxon: scientificName: Telothyriamanuelpereirai; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Telothyria; specificEpithet: manuelpereirai; scientificNameAuthorship: Fleming & Wood, 2018; Location: continent: Central America; country: Costa Rica ; countryCode: CR; stateProvince: Alajuela; county: Sector Rincon Rain Forest; locality: Area de Conservacion Guanacaste ; verbatimLocality: Sendero Rincon; verbatimElevation: 430 ; verbatimLatitude: 10.8962; verbatimLongitude: -85.2777; verbatimCoordinateSystem: Decimal; decimalLatitude: 10.8962 ; decimalLongitude: -85.2777 ; Identification: identifiedBy: AJ Fleming ; dateIdentified: 2018; Event: samplingProtocol: Reared from the larve of the Crambidae, Piletosoma thialisDHJ03 ; verbatimEventDate: 13-Aug-2006 ; Record Level: language: en; institutionCode: CNC ; collectionCode: Insects ; basisOfRecord: Pinned Specimen Type status: Paratype . Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0011562 ; recordedBy: D.H. Janzen, W. Hallwachs & Minor Carmona ; individualID: DHJPAR0011562; individualCount: 1 ; sex: M ; lifeStage: adult ; preparations: pinned; otherCatalogNumbers: ASTAQ949-06, 05-SRNP-42657, BOLD:AAB8712; Taxon: scientificName: Telothyriamanuelpereirai; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Telothyria; specificEpithet: manuelpereirai; scientificNameAuthorship: Fleming & Wood, 2018; Location: continent: Central America; country: Costa Rica ; countryCode: CR; stateProvince: Alajuela; county: Sector Rincon Rain Forest; locality: Area de Conservacion Guanacaste ; verbatimLocality: Sendero Tucan; verbatimElevation: 410 ; verbatimLatitude: 10.9042; verbatimLongitude: -85.2712; verbatimCoordinateSystem: Decimal; decimalLatitude: 10.9042 ; decimalLongitude: -85.2712 ; Identification: identifiedBy: AJ Fleming ; dateIdentified: 2018; Event: samplingProtocol: Reared from the larve of the Crambidae, Piletosoma thialisDHJ03 ; verbatimEventDate: 15-Oct-2005 ; Record Level: language: en; institutionCode: CNC ; collectionCode: Insects ; basisOfRecord: Pinned Specimen Type status: Paratype . Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0011565 ; recordedBy: D.H. Janzen, W. Hallwachs & Minor Carmona ; individualID: DHJPAR0011565; individualCount: 1 ; sex: F ; lifeStage: adult ; preparations: pinned; otherCatalogNumbers: ASTAQ952-06, 05-SRNP-42654, BOLD:AAB8712; Taxon: scientificName: Telothyriamanuelpereirai; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Telothyria; specificEpithet: manuelpereirai; scientificNameAuthorship: Fleming & Wood, 2018; Location: continent: Central America; country: Costa Rica ; countryCode: CR; stateProvince: Alajuela; county: Sector Rincon Rain Forest; locality: Area de Conservacion Guanacaste ; verbatimLocality: Sendero Tucan; verbatimElevation: 410 ; verbatimLatitude: 10.9042; verbatimLongitude: -85.2712; verbatimCoordinateSystem: Decimal; decimalLatitude: 10.9042 ; decimalLongitude: -85.2712 ; Identification: identifiedBy: AJ Fleming ; dateIdentified: 2018; Event: samplingProtocol: Reared from the larve of the Crambidae, Piletosoma thialisDHJ03 ; verbatimEventDate: 12-Oct-2005 ; Record Level: language: en; institutionCode: CNC ; collectionCode: Insects ; basisOfRecord: Pinned Specimen Type status: Paratype . Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0011566 ; recordedBy: D.H. Janzen, W. Hallwachs & Minor Carmona ; individualID: DHJPAR0011566; individualCount: 1 ; sex: M ; lifeStage: adult ; preparations: pinned; otherCatalogNumbers: ASTAQ953-06, 05-SRNP-42649, BOLD:AAB8712; Taxon: scientificName: Telothyriamanuelpereirai; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Telothyria; specificEpithet: manuelpereirai; scientificNameAuthorship: Fleming & Wood, 2018; Location: continent: Central America; country: Costa Rica ; countryCode: CR; stateProvince: Alajuela; county: Sector Rincon Rain Forest; locality: Area de Conservacion Guanacaste ; verbatimLocality: Sendero Tucan; verbatimElevation: 410 ; verbatimLatitude: 10.9042; verbatimLongitude: -85.2712; verbatimCoordinateSystem: Decimal; decimalLatitude: 10.9042 ; decimalLongitude: -85.2712 ; Identification: identifiedBy: AJ Fleming ; dateIdentified: 2018; Event: samplingProtocol: Reared from the larve of the Crambidae, Piletosoma thialisDHJ03 ; verbatimEventDate: 15-Oct-2005 ; Record Level: language: en; institutionCode: CNC ; collectionCode: Insects ; basisOfRecord: Pinned Specimen Type status: Paratype . Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0006598 ; recordedBy: D.H. Janzen, W. Hallwachs & Jose Perez ; individualID: DHJPAR0006598; individualCount: 1 ; sex: M ; lifeStage: adult ; preparations: pinned; otherCatalogNumbers: ASTA777-06, 05-SRNP-43018, BOLD:AAB8712; Taxon: scientificName: Telothyriamanuelpereirai; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Telothyria; specificEpithet: manuelpereirai; scientificNameAuthorship: Fleming & Wood, 2018; Location: continent: Central America; country: Costa Rica ; countryCode: CR; stateProvince: Alajuela; county: Sector Rincon Rain Forest; locality: Area de Conservacion Guanacaste ; verbatimLocality: Sendero Rincon; verbatimElevation: 430 ; verbatimLatitude: 10.8962; verbatimLongitude: -85.2777; verbatimCoordinateSystem: Decimal; decimalLatitude: 10.8962 ; decimalLongitude: -85.2777 ; Identification: identifiedBy: AJ Fleming ; dateIdentified: 2018; Event: samplingProtocol: Reared from the larve of the Crambidae, Piletosoma thialisDHJ03 ; verbatimEventDate: 17-Nov-2005 ; Record Level: language: en; institutionCode: CNC ; collectionCode: Insects ; basisOfRecord: Pinned Specimen Type status: Paratype . Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0018609 ; recordedBy: D.H. Janzen, W. Hallwachs & Jose Perez ; individualID: DHJPAR0018609; individualCount: 1 ; sex: F ; lifeStage: adult ; preparations: pinned; otherCatalogNumbers: ASTAI1256-07, 02-SRNP-7398, BOLD:AAB8712; Taxon: scientificName: Telothyriamanuelpereirai; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Telothyria; specificEpithet: manuelpereirai; scientificNameAuthorship: Fleming & Wood, 2018; Location: continent: Central America; country: Costa Rica ; countryCode: CR; stateProvince: Alajuela; county: Sector Rincon Rain Forest; locality: Area de Conservacion Guanacaste ; verbatimLocality: Sendero Rincon; verbatimElevation: 430 ; verbatimLatitude: 10.8962; verbatimLongitude: -85.2777; verbatimCoordinateSystem: Decimal; decimalLatitude: 10.8962 ; decimalLongitude: -85.2777 ; Identification: identifiedBy: AJ Fleming ; dateIdentified: 2018; Event: samplingProtocol: Reared from the larve of the Crambidae, Piletosoma thialisDHJ02 ; verbatimEventDate: 03-Jul-2002 ; Record Level: language: en; institutionCode: CNC ; collectionCode: Insects ; basisOfRecord: Pinned Specimen Type status: Paratype . Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0018610 ; recordedBy: D.H. Janzen, W. Hallwachs & Jose Perez ; individualID: DHJPAR0018610; individualCount: 1 ; sex: F ; lifeStage: adult ; preparations: pinned; otherCatalogNumbers: ASTAI1257-07, 02-SRNP-7399, BOLD:AAB8712; Taxon: scientificName: Telothyriamanuelpereirai; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Telothyria; specificEpithet: manuelpereirai; scientificNameAuthorship: Fleming & Wood, 2018; Location: continent: Central America; country: Costa Rica ; countryCode: CR; stateProvince: Alajuela; county: Sector Rincon Rain Forest; locality: Area de Conservacion Guanacaste ; verbatimLocality: Sendero Rincon; verbatimElevation: 430 ; verbatimLatitude: 10.8962; verbatimLongitude: -85.2777; verbatimCoordinateSystem: Decimal; decimalLatitude: 10.8962 ; decimalLongitude: -85.2777 ; Identification: identifiedBy: AJ Fleming ; dateIdentified: 2018; Event: samplingProtocol: Reared from the larve of the Crambidae, Piletosoma thialisDHJ02 ; verbatimEventDate: 04-Jul-2002 ; Record Level: language: en; institutionCode: CNC ; collectionCode: Insects ; basisOfRecord: Pinned Specimen Type status: Paratype . Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0018611 ; recordedBy: D.H. Janzen, W. Hallwachs & Jose Perez ; individualID: DHJPAR0018611; individualCount: 1 ; sex: F ; lifeStage: adult ; preparations: pinned; otherCatalogNumbers: ASTAI1258-07, 02-SRNP-7401, BOLD:AAB8712; Taxon: scientificName: Telothyriamanuelpereirai; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Telothyria; specificEpithet: manuelpereirai; scientificNameAuthorship: Fleming & Wood, 2018; Location: continent: Central America; country: Costa Rica ; countryCode: CR; stateProvince: Alajuela; county: Sector Rincon Rain Forest; locality: Area de Conservacion Guanacaste ; verbatimLocality: Sendero Rincon; verbatimElevation: 430 ; verbatimLatitude: 10.8962; verbatimLongitude: -85.2777; verbatimCoordinateSystem: Decimal; decimalLatitude: 10.8962 ; decimalLongitude: -85.2777 ; Identification: identifiedBy: AJ Fleming ; dateIdentified: 2018; Event: samplingProtocol: Reared from the larve of the Crambidae, Piletosoma thialisDHJ02 ; verbatimEventDate: 03-Jul-2002 ; Record Level: language: en; institutionCode: CNC ; collectionCode: Insects ; basisOfRecord: Pinned Specimen Type status: Paratype . Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0035703 ; recordedBy: D.H. Janzen, W. Hallwachs & Jose Perez ; individualID: DHJPAR0035703; individualCount: 1 ; sex: M ; lifeStage: adult ; preparations: pinned; otherCatalogNumbers: ASHYD1084-09, 09-SRNP-41306, BOLD:AAB8712; Taxon: scientificName: Telothyriamanuelpereirai; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Telothyria; specificEpithet: manuelpereirai; scientificNameAuthorship: Fleming & Wood, 2018; Location: continent: Central America; country: Costa Rica ; countryCode: CR; stateProvince: Alajuela; county: Sector Rincon Rain Forest; locality: Area de Conservacion Guanacaste ; verbatimLocality: Estacion Caribe; verbatimElevation: 415 ; verbatimLatitude: 10.9019; verbatimLongitude: -85.2749; verbatimCoordinateSystem: Decimal; decimalLatitude: 10.9019 ; decimalLongitude: -85.2749 ; Identification: identifiedBy: AJ Fleming ; dateIdentified: 2018; Event: samplingProtocol: Reared from the larve of the Crambidae, Piletosomathialis ; verbatimEventDate: 14-Jul-2009 ; Record Level: language: en; institutionCode: CNC ; collectionCode: Insects ; basisOfRecord: Pinned Specimen Type status: Paratype . Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0035816 ; recordedBy: D.H. Janzen, W. Hallwachs & Noe Castillo ; individualID: DHJPAR0035816; individualCount: 1 ; sex: F ; lifeStage: adult ; preparations: pinned; otherCatalogNumbers: ASHYD1197-09, 09-SRNP-69212, BOLD:AAB8712; Taxon: scientificName: Telothyriamanuelpereirai; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Telothyria; specificEpithet: manuelpereirai; scientificNameAuthorship: Fleming & Wood, 2018; Location: continent: Central America; country: Costa Rica ; countryCode: CR; stateProvince: Alajuela; county: Sector Rincon Rain Forest; locality: Area de Conservacion Guanacaste ; verbatimLocality: Jacobo; verbatimElevation: 461 ; verbatimLatitude: 10.9408; verbatimLongitude: -85.3177; verbatimCoordinateSystem: Decimal; decimalLatitude: 10.9408 ; decimalLongitude: -85.3177 ; Identification: identifiedBy: AJ Fleming ; dateIdentified: 2018; Event: samplingProtocol: Reared from the larve of the Crambidae, Piletosoma thialisDHJ02 ; verbatimEventDate: 19-Jul-2009 ; Record Level: language: en; institutionCode: CNC ; collectionCode: Insects ; basisOfRecord: Pinned Specimen Type status: Paratype . Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0035819 ; recordedBy: D.H. Janzen, W. Hallwachs & Noe Castillo ; individualID: DHJPAR0035819; individualCount: 1 ; sex: F ; lifeStage: adult ; preparations: pinned; otherCatalogNumbers: ASHYD1200-09, 09-SRNP-69246, BOLD:AAB8712; Taxon: scientificName: Telothyriamanuelpereirai; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Telothyria; specificEpithet: manuelpereirai; scientificNameAuthorship: Fleming & Wood, 2018; Location: continent: Central America; country: Costa Rica ; countryCode: CR; stateProvince: Alajuela; county: Sector Rincon Rain Forest; locality: Area de Conservacion Guanacaste ; verbatimLocality: Jacobo; verbatimElevation: 461 ; verbatimLatitude: 10.9408; verbatimLongitude: -85.3177; verbatimCoordinateSystem: Decimal; decimalLatitude: 10.9408 ; decimalLongitude: -85.3177 ; Identification: identifiedBy: AJ Fleming ; dateIdentified: 2018; Event: samplingProtocol: Reared from the larve of the Crambidae, Piletosoma thialisDHJ03 ; verbatimEventDate: 18-Jul-2009 ; Record Level: language: en; institutionCode: CNC ; collectionCode: Insects ; basisOfRecord: Pinned Specimen Type status: Paratype . Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0040183 ; recordedBy: D.H. Janzen, W. Hallwachs & Ricardo Calero ; individualID: DHJPAR0040183; individualCount: 1 ; sex: M ; lifeStage: adult ; preparations: pinned; otherCatalogNumbers: ASHYE2350-11, 10-SRNP-71645, BOLD:AAB8712; Taxon: scientificName: Telothyriamanuelpereirai; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Telothyria; specificEpithet: manuelpereirai; scientificNameAuthorship: Fleming & Wood, 2018; Location: continent: Central America; country: Costa Rica ; countryCode: CR; stateProvince: Guanacaste; county: Sector Pitilla; locality: Area de Conservacion Guanacaste ; verbatimLocality: Charia; verbatimElevation: 530 ; verbatimLatitude: 10.9934; verbatimLongitude: -85.4027; verbatimCoordinateSystem: Decimal; decimalLatitude: 10.9934 ; decimalLongitude: -85.4027 ; Identification: identifiedBy: AJ Fleming ; dateIdentified: 2018; Event: samplingProtocol: Reared from the larve of the Crambidae, Piletosomathialis ; verbatimEventDate: 09-Jul-2010 ; Record Level: language: en; institutionCode: CNC ; collectionCode: Insects ; basisOfRecord: Pinned Specimen Type status: Paratype . Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0040201 ; recordedBy: D.H. Janzen, W. Hallwachs & Ricardo Calero ; individualID: DHJPAR0040201; individualCount: 1 ; sex: M ; lifeStage: adult ; preparations: pinned; otherCatalogNumbers: ASHYE2368-11, 10-SRNP-71639, BOLD:AAB8712; Taxon: scientificName: Telothyriamanuelpereirai; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Telothyria; specificEpithet: manuelpereirai; scientificNameAuthorship: Fleming & Wood, 2018; Location: continent: Central America; country: Costa Rica ; countryCode: CR; stateProvince: Guanacaste; county: Sector Pitilla; locality: Area de Conservacion Guanacaste ; verbatimLocality: Charia; verbatimElevation: 530 ; verbatimLatitude: 10.9934; verbatimLongitude: -85.4027; verbatimCoordinateSystem: Decimal; decimalLatitude: 10.9934 ; decimalLongitude: -85.4027 ; Identification: identifiedBy: AJ Fleming ; dateIdentified: 2018; Event: samplingProtocol: Reared from the larve of the Crambidae, Piletosomathialis ; verbatimEventDate: 27-Jun-2010 ; Record Level: language: en; institutionCode: CNC ; collectionCode: Insects ; basisOfRecord: Pinned Specimen Description Male. Length: 7-10 mm (Fig. 34 ). Head (Fig. 34 b ): frons narrow, 1/5 of head width; gena 1/8 of head height; three reclinate orbital setae; anteriormost reclinate orbital subequal to uppermost frontal seta; ocellar setae present but so small as to appear absent; outer vertical seta absent; fronto-orbital plate over 90% silver, with brassy tinge at most covering ocellar triangle and adjacent area; fronto-orbital plate with short blonde hairs interspersed among frontal setae; parafacial pale silver; facial ridge bare; palpus with a slight spatulate club at apex, sparsely haired along outer margin; arista orange-brown smoothly tapered, minutely plumose microtrichia at most as long as width of arista; postpedicel orange over 50% of surface; postocular region behind margin of eye upper 1/3 gold, with lower 2/3 including gena silver tomentose; upper 1/3 of occiput gold tomentose. Thorax (Fig. 34 a , c ): golden tomentose, with four distinct dorsal stripes; entire thorax covered in dense plumose blonde hairs, hairs extending to cover scutellum; chaetotaxy: 5-6 postpronotal setae, basal setae arranged in a straight line; supra-alar setae 2:3; intra-alar setae 2:3; dorsocentral setae 3:3; acrostichal setae 3:3; katepisternum with two setae. Scutellum golden tomentose; two pairs of strong marginal setae (basal and subapical) and a small pair of crossed apical scutellar setae 1/8-1/10th as long as subapical scutellars; basal scutellar setae subequal in length to subapical setae; subapical setae straight; underside of scutellum bearing plumose blonde hairs below basal scutellar setae. Legs : foreleg with yellow ground color; midleg yellow coxa and femur with brown tibia and tarsal segments; hindleg yellow ground color on proximal 50% of femur, transitioning to dark brown distally, tibia and tarsal segments; anterior leg tibia with regular fringe of equally spaced setae along anteroventral surface, and two widely spaced posterodorsal setae. Wings : basicosta bright ivory white; all veins bare, with only 1-2 setulae at base of R4+5; calypters pale white translucent with narrow beige fringes. Abdomen (Fig. 34 a , c ): ground color yellow-orange; ST1+2 yellow over more than 50% of tergite, with dark brown ground color medially along the middorsal depression, extending into a longitudinal middorsal brown stripe up to posterior edge of T4; ST1+2 with golden plumose hairs present on ventral surface; T3 and T4 each with dense gold tomentum along anterior marginal 10%, thinning and extending over remainder of tergite; T5 entirely orange with gold tomentum; one pair of median marginal setae weak almost undifferentiated on T3; row of marginal setae present only on T4 and T5; median discal setae absent. Male terminalia (Fig. 34 d , e , f ): Sternite 5 with a wide deeply excavated median cleft, smoothly V-shaped, margins covered in dense pollinosity; lateral lobes of sternite pointed apically, with a small group of strong setulae along outer margins; basal section of sternite 5 subequal to slightly shorter than length of apical lobes. Cerci in posterior view sharply pointed and slender, abruptly widening to a moderate almost rectangular shoulder along the basal section, equal in length to surstyli, fused along entire length; in lateral view, with a smooth regular downward curve along apical 2/3rds; several strong widely spaced setulae along basal 1/3rd. Surstylus in lateral view, almost equilateral along its length rounded at tip, digitiform; surstylus appearing fused with epandrium; when viewed dorsally surstyli appear slender with an inward bend. Distiphallus 2X as long as basiphallus and tubular, slightly pointed at apex. Female. Length: 5-9 mm (Fig. 35 ). Head (Fig. 35 b ): as in male with the following exceptions: fronto-orbital plate 90% silver; parafacial brilliant silver; frons 1/5 of head width; three inner reclinate orbital setae; two proclinate orbital setae; and outer vertical setae present. Thorax (Fig. 35 a , c ): katepisternum with three setae; meron densely covered in plumose hairs as in male but with the addition of 2-4 typical meral setae. Legs: colored as in male. Abdomen (Fig. 35 a , c ): ground color brown dorsally on ST1+2 and T3 with large patches of orange ventrolaterally; T4 entirely brown ground color; T5 entirely orange with gold tomentum; T3 with one strong pair of median marginal setae and T4, T5 with a complete row of marginal setae; T4 bearing two weak discal setae scarcely distinguishable from surrounding hairs. Diagnosis Telothyria manuelpereirai sp. n. can be distinguished from all other Telothyria by the following combination of traits: ocellar setae present but so small as to appear absent, postpedicel orange over 50% of surface, entire thorax covered in dense plumose blonde hairs, hairs extending to cover scutellum, katepisternum with two setae in males and three setae in females, and T5 yellow with silver tomentum. Etymology Telothyria manuelpereirai sp. n. is named in recognition of Manuel Pereira's outstanding work on the team that conducts the caterpillar and parasite inventory from ACG's Estacion Biologica Cacao. Distribution Costa Rica, ACG, Alajuela and Guanacaste Provinces, 410-530 m elevation. Ecology Telothyria manuelpereirai sp. n. has been reared 20 times from three species of Lepidoptera in the family Crambidae : Piletosoma thialis Dyar, 1914, Piletosoma thialis DHJ02, and Piletosoma thialis DHJ03, in rain forest.