<document id="D8D39CBB8E5B3BA469984A7C31BA5E53" ID-CLB-Dataset="20208" ID-DOI="10.11646/zootaxa.5115.4.1" ID-GBIF-Dataset="b99d2dc5-e0a6-4f73-b2c6-b21b6350c69f" ID-ISSN="1175-5326" ID-Zenodo-Dep="6361552" ID-ZooBank="320C2E00-8035-431A-8AD7-D483D81923F6" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1647419732391" checkinUser="plazi" docAuthor="Ullah, Mishkat, Naeem, Muhammad, Mahmood, Khalid &amp; Garner, Beulah" docDate="2022" docId="291CD3215311DC7BFF6DFE4C87D3FD14" docLanguage="en" docName="zootaxa.5115.4.1.pdf" docOrigin="Zootaxa 5115 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Chlaenius Bonelli 1810" docType="treatment" docVersion="7" lastPageNumber="457" masterDocId="D525AB595317DC7DFFFAFFFA836DFF91" masterDocTitle="Addition to the knowledge of Tribe Chlaeniini Brullé, 1834 (Coleoptera: Carabidae) from Pakistan" masterLastPageNumber="486" masterPageNumber="451" pageNumber="457" updateTime="1699318106451" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="96BC771695593A746432BF7785B92062">Addition to the knowledge of Tribe Chlaeniini Brullé, 1834 (Coleoptera: Carabidae) from Pakistan</mods:title>
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<mods:namePart id="AAB7DFCECE8EE7B201D737EE1AA4D900">Ullah, Mishkat</mods:namePart>
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<mods:namePart id="216E1FA52C7C491C435A071263715BDC">Naeem, Muhammad</mods:namePart>
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<mods:namePart id="F60017245A6F2D7D48097341230DAAB6">Mahmood, Khalid</mods:namePart>
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<mods:namePart id="8E04A07DCC11926E33F630E4EC2BFD14">Garner, Beulah</mods:namePart>
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<mods:affiliation id="8574100C41DD82F4A0C21515CCC7532B">Insects Division, Department of Life Sciences, Natural History Museum, Cromwell Road, London SW 7 5 BD, United Kingdom. &amp; b. garner @ nhm. ac. uk; https: // orcid. org / 0000 - 0002 - 5229 - 2450</mods:affiliation>
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Genus 
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<bibRefCitation id="C5241FC65311DC7BFE88FE4C817EFE41" author="Bonelli, F. A." box="[370,531,438,464]" pageId="6" pageNumber="457" pagination="1 - 58" refId="ref25707" refString="Bonelli, F. A. (1810) Observations entomologiques. Premiere partie (cicindeletes et portion des carabiques) [with the &quot; Tabula synoptica exhibens genera carabicorum in sectiones et stirpes disposita &quot;]. Memoires de l'Academie des Sciences, Turin, XVIII, 1 - 58." type="journal article" year="1810">Bonelli, 1810</bibRefCitation>
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The genus 
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<emphasis id="93C1BE255311DC7BFEEEFE0582EBFD88" box="[276,390,511,537]" italics="true" pageId="6" pageNumber="457">Chlaenius</emphasis>
Bonelli
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is the most species rich genera of the tribe 
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Chlaeniini 
<bibRefCitation id="C5241FC65311DC7BFBB4FE0587B1FD88" author="Brulle, G. A." box="[1102,1244,511,537]" pageId="6" pageNumber="457" refId="ref25801" refString="Brulle, G. A. (1834) Histoire naturelle des insectes, traitant de leur organisation et de leurs moeurs en general, par M. V. Audouin, et comprenant leur classification et la description des especes, par M. A. Brulle. Le tout accompagne de planches gravees sur acier, d'apres des peintures executees pour cette edition sur la collection du Museum de Paris. Tome IV. Coleopteres I. F. D. Pillot, Paris, 240 pp., 8 pls. [DP: 2 August 1834 (Bibl. Fr.), Note: This volume contains viii + 479 pages + 16 plates and was issued in two livraisons, the first one in 1834 and the second one in 1835. Despite the title, Brulle alone was the author]" type="book" year="1834">Brullé, 1834</bibRefCitation>
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with 855 known species worldwide (
<bibRefCitation id="C5241FC65311DC7BFE8CFDD98164FDAC" author="Lorenz, W." box="[374,521,546,573]" pageId="6" pageNumber="457" refId="ref27271" refString="Lorenz, W. (2005) A systematic list of extant ground beetle of the World. (Insecta, Coleoptera, Adephaga: Trachypachidae &amp; Carabidae incul. Paussinae, Cicindelinae, Rhysodinae). 2 nd Edition. Lorenz Publishing, Tutzing, 530 pp." type="book" year="2005">Lorenz 2005</bibRefCitation>
) arranged in 62 subgenera. It is widely distributed in all zoogeographical regions of the World, however, is more diverse in the Afrotropical and Oriental regions with approximately 650 species (
<bibRefCitation id="C5241FC65311DC7BFF65FD918223FD14" author="Bousquet, Y." box="[159,334,619,645]" pageId="6" pageNumber="457" pagination="1 - 1722" refId="ref25759" refString="Bousquet, Y. (2012) Catalogue of Geadephaga (Coleoptera, Adephaga) of America, North of Mexico. ZooKeys, 245, 1 - 1722. https: // doi. org / 10.3897 / zookeys. 245.3416" type="journal article" year="2012">Bousquet 2012</bibRefCitation>
). In 
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, itis represented by 29 species and 2 subspecies in 13 subgenera.
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