Phylogenetic systematics of the genera of Thryptocerina Jeannel, 1949 and new species from New Caledonia (Coleoptera, Carabidae, Oodini) Author Will, Kipling https://orcid.org/0000-0002-7056-9011 University of California Berkeley, Essig Museum of Entomology. Berkeley, California, USA kipwill@berkeley.edu Author Gueorguiev, Borislav National Museum of Natural History, 1 Blvd. Tsar Osvoboditel, 1000 Sofia, Bulgaria text ZooKeys 2021 2021-06-16 1044 375 425 http://dx.doi.org/10.3897/zookeys.1044.63775 journal article http://dx.doi.org/10.3897/zookeys.1044.63775 1313-2970-1044-375 D50CC77C2E6941FDA9BD395B025C43AE EBD9E81E5EC55AA491952682787793A4 Adelopomorpha glabra Heller, 1916 Figs 14 , 29 , 39 , 42 , 43 , 52 Adelopomorpha glabra Heller, 1916: 270. Material examined. Holotype : New Caledonia • ♀ ; card-mounted teneral specimen, with original labels, " Drs. F. Sarasin & J. Roux Neukaledonien Panie Wald" // "Adelop. glabra Typus" // "1914" // "Staatl. Mus. fuer Tierkunde Dresden" // "Senkenberg Naturhist. Sammlungen Dresden, Museum fuer Tierkunde (MTD) Museum fuer Tierkunde Dresden (MTD) MTD-COL-Car-Type-00921" [MTKD]. Type locality. Mt. Panie . Other material. New Caledonia • ♂ ; Mt. Panie ; 950-1300 m ; 14-16.v.1984 ; G. Monteith & D. Cook ; [QM] • ♀ ; same data as for preceding; [QM] • ♂ ; 20°34'S , 164°46'E ; Mt. Panie track; 1500 m ; 9.xi.2001 ; Pyrethrum , trees & logs; C. Burwell ; [QM] . Diagnosis. A glabra group species with depressed lateral 1/3 of the pronotum present (Fig. 43 ). See diagnosis of A. tuberculata and A. tethys for characteristics to distinguish those very similar species. The aedeagus is decisively different (Fig. 29 ), with a relatively long and broadly rounded tip, and with the membranous region of the ostium nearly reaching the apex. Redescription. Habitus . Small sized, BL: 5.69 mm (range, BL: 5.69-6.00 mm), BW: 2.89 mm (range, 2.83-2.89 mm). Color and luster . Head and pronotum dark reddish brown to nearly black on pronotal disc; elytra and ventral surface of body black, pronotum marginally diaphanous reddish brown; legs deep reddish brown; Ventrite 6 reddish brown, ventrite 5 black or medially at the very apical edge reddish black; antennae and palpi brown. Integument moderately dull, with diffused spectral iridescence dorsally and ventrally, more prominent on elytra. Microsculpture and punctation . Dorsal surface of head and pronotum with prominent isodiametric meshes; elytra with somewhat transverse microsculpture; ventral surface with scarcely visible sculpticells or sculpticells mostly not apparent, most apparent on prosternal process. Head lacking macropunctation or wrinkles on vertex; head, pronotum and elytra with extremely fine, scattered micropunctation or lacking such punctures; all abdominal ventrites very shallowly wrinkled laterally and smooth medially. Head . Approximately 1/3 as wide as pronotum (Suppl. material 2: Table S1). Eye moderate sized, somewhat to moderately protruded, EyW/HW: 1.26 (range, 1.26-1.41; the low value in the type may be due to head position in image). Frontoclypeal sutures not evident. Mentum tooth without paramedial border. Thorax . Pronotum 1 2/3 wider than long (PW/PL: 1.70 (range, 1.60-1.70)), with sides evenly, very shallowly rounded from anterior angles to posterior ones; maximal width in posterior 1/5; width at apex ~ 1/3 that of widest point (PW/PA: 2.60 (range 2.32-2.60)). Posterior angles ~ right angled, rounded. Mesosternum with single, low medial tubercle. Metepisternum 1 1/3 x wider than long, with lateral margin slightly convex, coadunation with epipleuron along entire length. Elytra . Broad, as wide as long (EL/EW: 1.06 (range, 1.11-1.15)). Basal margin distinct, forming very sharp tooth at shoulder, ended medially at level of parascutellar punctures, joined to parascutellar puncture fovea by a short, shallow striole. Apical sinuation not evident. Female genitalia . Bursa copulatrix small in comparison with spermatheca; spermatheca narrow, undifferentiated, coiled at distal 2/3; spermathecal gland connected near basal 1/5-1/4 of spermatheca (Fig. 14 ). Male genitalia . Median lobe of aedeagus in lateral view long, shallowly curved (Fig. 39 ); lobe in dorsal view, with apical lamella, short, rounded, and very slightly oriented to right (Fig. 29 ); sclerotized portion of endophallus visible in repose, with two sclerites.