Pipunculidae (Diptera) from southern Iran, including two new species of the genus Tomosvaryella Aczél Author Majnon-Jahromi, Bahareh Author Gheibi, Mehdi Author Fallahzadeh, Majid Author Kehlmaier, Christian Author Hesami, Shahram text Zootaxa 2017 4273 4 488 500 journal article 32892 10.11646/zootaxa.4273.4.2 316e1968-2a32-462e-a554-99165078f1fc 1175-5326 803688 77091F8E-2A3C-4A9B-81C2-656B5BF72C22 Tomosvaryella angulata Kehlmaier & Majnon-Jahromi sp. nov. Figs 3–8 Type material. HOLOTYPE: 1♂, loc. 7, 9–16.v.2014 (SMTD); PARATYPES: 1♂ (DNA CK831, LT671787 ), loc. 7, 9–16.v.2014 (SMTD); 1♂, loc. 4, 18–28.vi.2015 (SHIAU); 1♀ (DNA CK804, LT671786 ), loc. 3, 5–15.iv.2012 (SMTD). Description. MALE . Body length 2.9–3.3 mm ; wing length 3.5–3.8 mm ; eyes meeting for a distance slightly more than twice the length of ocellar triangle, i.e., for approximately six to nine ommatidia; pedicellus with two minute bristles dorsally and one ventrally; flagellum pointed, tawny, tip may be paler; thorax dark brown except postpronotal lobe yellow; dorsal bristles hardly discernible, black; wing membrane with a faint brownish tinge; wing membrane covered in microtrichia except for the base, including the anal lobe basally; third costal segment 1/ 3 the length of fourth costal segment; crossvein r-m reaching cell dm at or shortly after the middle; crossvein bm-cu interrupted right before reaching cell br on both wings; halter with dark base, whitish stem and light brown knob; leg dark brown except for the narrowly yellow apex of femora, basal 1/3 and apex of tibia and tarsal segments except distitarsus; hind trochanter ventrally smooth, without spines or projection; fore femora with posteroventral row of minute black spines in apical half; mid femora with antero- and posteroventral row of minute black spines for more than apical half; hind femora with anteroventral row of minute black spines in apical half; pulvilli slightly shorter than distitarsus; abdomen dark brown; tergite 1 with lateral patch of about ten short black setae on each side, slightly shorter than width of tarsal segments; dorsal setae of abdomen black, hardly discernible; sclerites dark brown, apparently evenly sclerotised; genitalia, in dorsal view, with short epandrium, broader than long, slightly less sclerotised in the central part; surstyli almost symmetrical, gradually tapering towards apices, which are distinctly knobbed; left surstylus slightly protruding the right one; membrane of syntergosternite 8 large, almost reaching epandrium, caudally occupying 1/3 of syntergosternite 8; in lateral view, apical half of surstyli bent towards venter by 90° (see inner margin); phallus trifid, with two longer and one shorter upright and wavy ejaculatory ducts, the longest one carrying about six scale-like projections at its base (see at high magnification, i.e., 100x); phallic guide small and inconspicuous, tip pointing towards epandrium. FEMALE . Body length 2.7 mm ; wing length 3.2 mm , differing from male by dichoptic eyes; frontal ommatidial facets distinctly enlarged; frons widened in its middle, twice the width of the ocellar triangle or equal to the length of four enlarged ommatidial facets; flagellum with paler apical half; crossvein r-m reaching cell dm shortly after the middle; pulvilli of fore and mid legs as long as distitarsi; ovipositor with dark base and yellow piercer; in lateral view, base of ovipositor with a distinct ventral protuberance; piercer gently undulating; in dorsal view, base of ovipositor rather trapezoid and piercer with weak lateral flanges in the middle of its length. Differential diagnosis. To the best of our knowledge, T. angulata sp . nov . is a unique species among the Palaearctic Pipunculidae , due to its distinctly apically-knobbed and laterally-bent surstyli in the male, and due to the weakly undulating ovipositor that bears a distinct ventral protuberance in the female ( Figs 3–8 ). For further remarks, see under T. pistacia sp . nov . Etymology. The Latin adjective angulata (English: angular) refers to the angled surstyli of male genitalia.