Revision of the genus Eulichas Jacobson, 1913 (Coleoptera: Eulichadidae) II. E. dudgeoni species group Author Hájek, Jiří text Zootaxa 2009 2192 1 44 journal article 10.5281/zenodo.189396 ea4e0f79-8de5-4af5-b12c-b88b00befe28 1175-5326 189396 Eulichas alesbezdeki Hájek , sp. nov. ( Figs. 1 , 25 , 45 , 65 ) Eulichas sp. (Guangxi A) cf. funebris et undulata : Jäch 1995 : 370 (partim). Eulichas jendeki : Jäch 1995 : 373 (partim). Type locality. Laos , Boli Kham Xai Prov., 8 km NE of Ban Nape. Type material. 79 specimens — Holotype 3 ( NMPC ), labelled: “LAOS-CE, 1–18.v.2001 , / Boli Kham Xai prov., / 18°21’N 105°08’E , / BAN NAPE ( 8 km NE ), / ~ 600m , Vít Kubáň leg. [printed] / Biological expedition / „ Laos 2001“ / Moravian Museum Brno / Czech Republic [printed]”. Paratypes : 2033 (nos. 1– 20) 6ƤƤ (nos. 21–26), same label data as holotype ( MZMB , NHMW , NMPC ); 1333 (nos. 27–39) 4ƤƤ (nos. 40–43), same label data, but “L. Dembický leg.” ( NHMB , NMPC ); 233 (nos. 44–45), “ LAOS – NE ; HUA PHAN prov. / 25km SE Vieng Xai (by road); / BAN KANGPABONG env.; / 20°19’N 104°25’E ; / J. Bezdĕk leg.; 14.–18.v.2001 [printed]” ( NMPC ); 13 (no. 46), “LAOS-C, Kamphéng Nakhon / Viangchan prov., Viang Chan / (=VIENTIANE), 17°58’N / 102°36’E , 170m , 17.v.2008 , / Khampaseuth Sisoutham leg. [printed]” ( NMPC ); 13 (no. 47), “ Tonkin / Than-Moi / Juni-Juli / H.Fruhstorfer [printed]” ( HNHM ); 633 (nos. 48–53), “ Tonkin / Montes Mauson [Tam Dảo], / April, Mai 2 -3000’ / H.Fruhstorfer [printed]” ( HNHM , ZMHB ); 233 (nos. 54–55), “ Vietnam Tam dao / Vinh phu pr. / 3.– 11.6.1985 / J. Picka lgt. [printed]” ( NHMB ); 1Ƥ (no. 56), “ Vietnam / Tam dao 3.6.– 11.6.1985 / Vinh phu prov. / Strnad Jan lgt. [printed]” ( NMPC ); 13 (no. 57), “ Vietnam N / Tam dao 26.5.– 3.6.1986 / Vinh phu prov. / Strnad Jan lgt. [printed]” ( NMPC ); 433 (nos. 58–61) 1Ƥ (no. 62), “N. VIETNAM 900m / TamDao 13.–24. / 5.1989 A. Olexa [printed]” ( NMPC ); 13 (no. 63) 3ƤƤ (nos. 64–66), “ VIETNAM 20.–28.VI.1990 / Vinh Phu Prov. / TAM DAO / Jan Strnad leg. [printed]” ( NMPC ); 1Ƥ (no. 67), “N. Vietnam / Tamdao / 20.6.1995 Gorochov [handwritten]” ( ZMAS ); 1Ƥ (no. 68), “N-Vietnam, Tam Dao, Vinh / Phu Prov., 21°27’18’’N ; / 105°38’58’’E / 2.–6.VI.1999 , 1050– 1200m ; / leg.: Fabrizi & Ahrens [printed]” ( SMTD ); 13 (no. 69), “ Vietnam gory [Mts.] SW / Cua-rao 600 m / 24.6.196 Kabakov [printed, transcribed from Cyrillic]” ( ZMAS ); 13 (no. 70), “ Vietnam / Lao Cai [Lào Cai, ca. 22°30’N 103°57’E ] / 15.5.1960 Izokh [handwritten, transcribed from Cyrillic]” ( ZMAS ); 13 (no. 71), “ Vietnam gory [Mts.] SW / Bai Tuong [Bái Thuơọơng] 300 m / 5.7.1962 Kabakov [printed, transcribed from Cyrillic]” ( ZMAS ); 1Ƥ (no. 72), “ Vietnam gory [Mts.] / W Sa-pa [Sa Pa, ca. 22°20’N 103°50’E ] 1600–2000 m / 1.6.1963 Kabakov [printed, transcribed from Cyrillic]” ( ZMAS ); 13 (no. 73), “ Vietnam gory [Mts.] 50 km / NO [ NE ] Thai Nguyen [ Thái Nguyên, ca. 21°33’N , 105°51’E ] 300 m / 15.6.1963 Kabakov [printed, transcribed from Cyrillic]” ( ZMAS ); 13 (no. 74), “ Vietnam gory [Mts.] 60 km / NO [ NE ] Thai Nguyen 300 m / 16.6.1963 Kabakov [printed, transcribed from Cyrillic]” ( ZMAS ); 1Ƥ (no. 75), “16.– 17.5.1990 YEN BAI [Yên Bái] / HOANG LIEN SON Distr. / N VIETNAM / JAN HORÁK Leg. [printed]” ( NMPC ); 233 (nos. 76–77) 1Ƥ (no. 78), “ China / Yunnan [printed]” ( HNHM , NMPC ). Additional material studied. 12 specimens — LAOS : 1Ƥ, Annam ( HNHM ). VIETNAM : 1Ƥ, H t Tonkin , A.N. Claire ( IRSNB ); 1Ƥ, Tonkin , Rég.[ion] Hà Giang, 1916, S. Olivier ( MNHN ); 1Ƥ, Tonkin , Chapo, prov. de Laokay [Sa Pa, ca. 22°20’N 103°50’E ; Lao Cai prov.] ( MNHN ); 1Ƥ, Tonkin , NE of Bảo Lạc [ca. 22°57’N 105°39’E ], 1897–1898 , Dr. Battarel leg. ( MNHN ); 1Ƥ, Làng Son [ca. 21°51’N 106°45’E ] ( MNHN ); 13, Tonkin , Than-Moi [most probably Thanh Moi, ca. 21°38’N 106°32’E ], v.–vi., H. Fruhstorfer ( MNHN ) [ paratype of E. jendeki ]; 13, Tonkin , Chiêm Hóa, viii.–ix., H. Fruhstorfer ( MNHN ) [ paratype of E. jendeki ]; 13, Tonkin , Hòa Bình env. [ca. 20°50’N 105°19’E ], 1902, J. Laisi leg. ( MNHN ) [ paratype of E. jendeki ]; 233, Tonkin , Rég.[ion] Hòa Bình, 1919 ( MNHN ) [ paratypes of E. jendeki ]; 13, Tonkin , Backan [Bảc Cạn, ca. 22°08’N 105°49’E ], 1907–08 , P. Lemée ( NHMW ) [ paratype of E. jendeki ]. Description. Habitus elongate, fusiform. Body colouring from brownish-red to black. Pale part of setation consists of recumbent yellowish or greyish setae forming typical ocellations on pronotum, elytra and abdominal sternites ( Fig. 1 ). Measurements. Males: 21–26 mm ( holotype 22 mm ); females: 25–31 mm . Head punctation consists of sparse moderately large setigerous punctures. Antenna robust, last antennomere nearly rectangular, 2.57–3.06 times as long as wide ( Fig. 25 ), its ventral side smooth. Pronotum trapezoidal, ca. 1.89–2.05 times as wide as long. Sides almost regularly rounded ( Fig. 45 ). The disc with two oval shallow depressions. Punctation consists of sparse moderately coarse setigerous punctures on the disc, which become slightly coarser and denser laterally. Elytra with numerous longitudinal rows of moderately large punctures, and very fine interstitial setigerous punctures. Ventral part with fine punctures, which are sparse medially and become larger and denser laterally. Last abdominal ventrite laterally regularly rounded to apex. Male. Aedeagus with phallobase shorter than parameres. Parameres simple, relatively slender, their subbasal hook small, but the subapical hook is well developed and prominent. Median lobe narrowly lanceolate ( Fig. 65 ). Female. Similar to male in habitus, but larger. Antenna shorter, and more slender. Differential diagnosis. The new species is similar to E. jendeki in habitus, while its aedeagus is rather close to that of E. dudgeoni . Eulichas alesbezdeki sp. nov. differs from E. jendeki in the almost regularly rounded sides of pronotum and the slender and slightly attenuating parameres of aedeagus, and from E. dudgeoni in shorter last antennomere and distinct body ocellations. FIGURES 1–12. Habitus of Eulichas . 1— E. alesbezdeki sp. nov. (holotype); 2— E. bertiae (paratype, Suisapa); 3— E. dudgeoni (paratype, Hong Kong); 4— E. jendeki (Sa Pa) ; 5— E. siamensis sp. nov. (paratype, Ban Bo Klua); 6— E. similis sp. nov. (paratype, Soppong); 7— E. uniformis (Dolakha) ; 8— E. baeri (Luzon) ; 9— E. baeri (Crocker Range) ; 10— E. gigantea (holotype); 11— E. incisicollis ♂ (Tanah Rata) ; 12— E. incisicollis Ƥ (holotype). FIGURES 13–24. Habitus of Eulichas . 13— E. oborili sp. nov. (holotype); 14— E. robusta sp. nov. (holotype); 15— E. sausai sp. nov. (holotype); 16— E. serricornis (paratype, Felda Lasah); 17— E. sikkimensis (Danda Pakhar) ; 18— E. sundaensis sp. nov. (holotype); 19— E. sundaensis sp. nov. Ƥ (Java); 20— E. wewalkai sp. nov. (holotype); 21— E. fasciolata (Crocker Range) ; 22— E. jakli sp. nov. (holotype); 23— E. subocellata (Tawau) ; 24— E. villosa sp. nov. (paratype, Gunung Emas). FIGURES 25–44. Male antennomeres IX–XI of Eulichas . 25— E. alesbezdeki sp. nov. ; 26— E. bertiae ; 27— E. dudgeoni ; 28— E. jendeki ; 29— E. siamensis sp. nov. ; 30— E. similis sp. nov. ; 31— E. uniformis ; 32— E. baeri ; 33— E. incisicollis ; 34— E. oborili sp. nov. ; 35— E. robusta sp. nov. ; 36— E. sausai sp. nov. ; 37— E. serricornis sp. nov. ; 38— E. sikkimensis ; 39— E. sundaensis sp. nov. ; 40— E. wewalkai sp. nov. ; 41— E. fasciolata ; 42— E. jakli sp. nov. ; 43— E. subocellata ; 44— E. villosa sp. nov. Scale bar 1 mm. FIGURES 45–54. Pronotum of Eulichas . 45— E. alesbezdeki sp. nov. ; 46— E. bertiae ; 47— E. dudgeoni ; 48— E. jendeki ; 49— E. siamensis sp. nov. ; 50— E. similis sp. nov. ; 51— E. uniformis ; 52— E. baeri (Luzon) ; 53— E. incisicollis ; 54— E. oborili sp. nov. Not in Scale. FIGURES 55–64. Pronotum of Eulichas . 55— E. robusta sp. nov. ; 56— E. sausai sp. nov. ; 57— E. serricornis sp. nov. ; 58— E. sikkimensis ; 59— E. sundaensis sp. nov. ; 60— E. wewalkai sp. nov. ; 61— E. fasciolata ; 62— E. jakli sp. nov. ; 63— E. subocellata ; 64— E. villosa sp. nov. Not in Scale. FIGURES 65–71. Aedeagus of Eulichas in dorsal view. 65— E. alesbezdeki sp. nov. ; 66— E. bertiae ; 67— E. dudgeoni ; 68— E. jendeki ; 69— E. siamensis sp. nov. ; 70— E. similis sp. nov. ; 71— E. uniformis . Scale bar 1 mm. FIGURES 72–77. Aedeagus of Eulichas in dorsal view. 72— E. baeri ; 73— E. incisicollis ; 74— E. oborili sp. nov. ; 75— E. robusta sp. nov. ; 76— E. sausai sp. nov. ; 77— E. serricornis sp. nov. Scale bar 1 mm. FIGURES 78–80. Aedeagus of Eulichas in dorsal view. 78— E. sikkimensis ; 79— E. sundaensis sp. nov. ; 80— E. wewalkai sp. nov. Scale bar 1 mm. Collection circumstances. Collected at light. Distribution. Known from several localities in north-eastern Laos , and “ Tonkin ”, northern Vietnam . Etymology. The new species is dedicated to my friend Aleš Bezdĕk (České Budĕjovice, Czech Republic ), specialist on Scarabaeoidea, with my special thanks for giving me the first Eulichas specimen and stimulating my interest in this family.