Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species Author Benedetti, Alipio Rezende Departamento de Zoologia, Instituto de Biociencias, Universidade de Sao Paulo, Caixa Postal: 11.461, 05422 - 970, Sao Paulo, SP, Brazil & Instituto Federal de Minas Gerais, campus Sao Joao Evangelista, Av. Primeiro de Junho, 1043, Sao Joao Evangelista, MG, Brazil alipiobenedetti@gmail.com Author Pinto-da-Rocha, Ricardo https://orcid.org/0000-0002-3959-2205 Departamento de Zoologia, Instituto de Biociencias, Universidade de Sao Paulo, Caixa Postal: 11.461, 05422 - 970, Sao Paulo, SP, Brazil text Arthropod Systematics & amp; Phylogeny 2022 2022-08-16 80 309 388 http://dx.doi.org/10.3897/asp.80.e73829 journal article http://dx.doi.org/10.3897/asp.80.e73829 1864-8312-80-309 D5C0468B99A14EF39237D9BC51A8BDA3 11907D1B6F665D96B5FD2F9751431366 3.52. Tschaidicancha scorsesei sp. nov. Figs 6E , 11K, L , 21D-F , 29A Description. MALE: Measurements ( n =10) DSW: 4.1-5.5 (5.5); DSL: 4.6-5.6 (5.6); CL: 1.6-2.2 (2.2). FIVL: 11.2-15.0 (12.4). ChL: 2.8-5.2 (5.2). Coloration : Brownish red DS. Chelicerae, pedipalpus and legs brown. Dorsum : (Fig. 6E ) Kappa-type DSS. Anterior margin of the carapace with median elevation; with sparse granules. Ocularium with pronounced median depression; with a pair of parallel spines. Carapace with few sparse granules. Areas I-IV with few sparse granules. Area I divided; with a medium pair of acuminate tubercles. Area III with a pair of spines, directed posteriorly, located in tegument elevations, very granular. Area IV short, with a pair of medium tubercles (smaller than those on Area I). Posterior margin of DS and free tergites I-III with a row of granules. Lateral margins of DS smooth. Chelicerae : (Fig. 6E ) Swollen on large males (as in the holotype). Slightly larger than female chelicerae in smaller males. Segment I slightly granular. Segment II with few granules; with five teeth. Segment III with two teeth. Pedipalpus : Trochanter with a ventroapical setiferous tubercle. Femur with a ventral row of 8-9 large tubercles (the middle ones largest) along the basal ⅔; one large prosubapical spine. Patella with a proapical tubercle. Tibia: retrolateral iIiIi, prolateral IiIi. Tarsus: retrolateral iiIiIi, prolateral iIIIi. Venter : Coxa I with a row of 4-5 tubercles. Coxae II-IV with granules throughout their surface. Genital area with few granules. Free sternites smooth. Anal operculum with few granules. Legs : (Figs 6E , 11K, L ) Coxae I-II each one with an anterior and a posterior apophysis. Coxa III unarmed. Coxa IV with sparse setiferous granules. Trochanters I-III unarmed and slightly granular. Trochanter IV with few granules and a retroapical small tubercle. Femora I-III unarmed and with small granules. Femur IV with sparse granules; a retrolateral row of 33-35 small acuminate tubercles, except at the apex. Patellae I-III unarmed, with few granules. Patella IV with three retroapical small tubercles. Tibiae I-III unarmed and with few granules. Tibia IV with a retrolateral row of 11-12 small acuminate tubercles along the basal 1/2 . Tarsal segmentation: ( n =10) 9-10 (10), 16-23 (23), 11-13 (12), 13-14 (14). Penis : (Fig. 21D-F ) VP subrectangular, long; distal margin straight; slightly curved in lateral view. MS C1-C2(3) subdistal long and slightly curved; MS A1 long and straight (slightly shorter than MS C and MS B), medially placed; MS B1 sub basal long and straight; MS D1 short and straight, placed below MS C. Lateral sacs long and apically blunt; with long T3-like microsetae. Stylus with several apical projections. Dorsal process present. Promontory convex. - FEMALE: Measurements ( n =5) DSW: 3.8-5.0; DSL: 4.3-5.6; CL: 1.5-2.2. FIVL: 10.5-15.3. ChL: 2.0-2.7. Chelicerae slightly smaller than that of small males and not swollen as in large males. Femur IV unarmed. Tarsal segmentation: ( n = 5) 7-9/15-17/11-14/12-14. Figure 31. Distribution of Metasarcus in Bolivia. A M. beni sp. nov. , M. bergmani sp. nov. , M. fellinii sp. nov. , M. limachii sp. nov. , M. vacafloresae ; B M. clavifemur , M. kurosawai sp. nov. , M. trispinosus sp. nov. Diagnosis. Similar to Tschaidicancha chaplini sp. nov. by the set of the following characteristics: reddish brown coloration; femur IV with a retrolateral row with more than 30 small tubercles; tibia IV with a retrolateral row with more than ten small tubercles (Fig. 11K, L ). Differs from T. chaplini sp. nov. by the set of the following characteristics: the area I divided longitudinally; free tergite III unarmed (Fig. 6E ); tibia IV with less than 15 retrolateral tubercles (Fig. 11K, L ); VP without distal-lateral projections (Fig. 21D-F ). Derivatio nominis. The specific epithet of masculine gender, in the genitive form, dedicated to the American director, producer, screenwriter and cinema historian Martin Charles Scorsese (1942-), an exponent of New Hollywood (American New Wave). Distribution. (Fig. 29A ) PERU. Pasco. Oxapampa. Parque Nacional Yanachaga-Chemillen . Material examined. Type material : Holotype ♂ , ' PERU , Pasco , Oxapampa , Parque Nacional Yanachaga-Chemillen , 10°32′42.1″S 75°21′24.4″W , 22/IV/2011 , R. Pinto-da-Rocha , A. Benedetti , J. Ochoa & D. Silva leg. (MUBI) - Paratypes 1 ♂ , 1 ♀ , 'ditto' (MZSP 36994). Additional material : 16 ♂ , 4 ♀ , ' PERU , Pasco , Oxapampa , Parque Nacional Yanachaga-Chemillen 10°32′42.1″S 75°21′24.4″W | 9-11/IV/2011 , S. Cardonel & J. Grados leg. (MUSM 0500505) .