A revision of Chrysopidae (Neuroptera) from the late Eocene Florissant Formation Colorado, with description of new species Author Makarkin, Vladimir N. Federal Scientific Center of the East Asia Terrestrial Biodiversity, Far East Branch of the Russian Academy of Sciences, Vladivostok, 960022, Russia Author Antell, Gwen S. Department of Earth Sciences, University of Oxford, Oxford OX 1 3 AN, United Kingdom Author Archibald, S. Bruce 0000-0002-4397-2497 Department of Biological Sciences, Simon Fraser University, 8888 University Drive, Burnaby, British Columbia, V 5 A 1 S 6, Canada sba 48 @ sfu. ca; https: // orcid. org / 0000 - 0002 - 4397 - 2497 & Museum of Comparative Zoology, 26 Oxford Street, Cambridge, Massachusetts, 02138, United States of America & Royal British Columbia Museum, 675 Belleville Street, Victoria, British Columbia, V 8 W 9 W 2, Canada sba48@sfu.ca text Zootaxa 2022 2022-05-05 5133 3 301 345 journal article 55712 10.11646/zootaxa.5133.3.1 dd1981ca-6b0b-42cf-839b-f1d3ba251c73 1175-5326 6522449 16A54EFD-028D-42FF-BD62-B8D35B7BA4DC Genus Dyspetochrysa Adams, 1967 Dyspetochrysa Adams, 1967: 219 , 220, 229; Schlüter 1984: 7; Willmann & Brooks 1991: 130 , 131; Adams & Penny 1992b: 216 ; Carpenter 1992: 347 ; Willmann 1993: 242 , 244; Peñalver et al . 1995: 481 , 485, 486; Makarkin 1998: 78 ; Nel et al. 2005: 67 ; Archibald et al . 2014: 191 ; Makarkin 2014: 4 ; Archibald & Makarkin 2015: 364 . Dyspectochrysa [ sic ]: Yang & Hong 1990: 20 . Type and only species. Tribochrysa vetuscula Scudder, 1890 , by original designation. Emended diagnosis. May be distinguished from other genera of Nothochrysinae by a combination of: Forewing: (1) M forked far distad origin of RP (shared with Archaeochrysa ) [other genera: proximad to slightly distad]; (2) one crossvein between RA, RP proximad 1r-m absent [ Archaeochrysa : present]; (3) im relatively narrow, elongate [ Cimbrochrysa , Danochrysa , Asiachrysa , Okanaganochrysa , Leptochrysa : broadly pentagonal; Hypochrysa , Asthenochrysa : broadly triangular]; (4) 2m-cu located in proximal part of im [ Cimbrochrysa , Asiachrysa : approximately in middle of im ; Pimachrysa , Pamochrysa , Kimachrysa , Hypochrysa , Asthenochrysa : proximad im ]. Remarks. The genus was defined by Adams (1967) based on the fusion of the forewing RP1 and MA for some distance in its only known specimen, a condition otherwise unknown in Nothochrysinae . However, this fusion occurs only in the right forewing ( Fig. 5B ) and these veins are separate in the left forewing ( Fig. 5C ). Therefore, we take this fusion to be an aberration, not diagnostic. In general, the preserved forewing venation of Dyspetochrysa is most similar to that of Archaeochrysa , especially by the far distal location of the M fork relative to the origin of RP. It is possible that these genera are synonyms, however, the crossvein in the RA space proximad 1r-m considered diagnostic of Archaeochrysa is absent in Dyspetochrysa , but this might also be an aberration. Further, the holotype and only known specimen of the only species of the genus, D. vetuscula is quite fragmentary. The validity of the genus is, therefore, tenuous. We prefer, however, to continue to recognize it here based on the diagnosis above, pending better preserved specimens to clarify its position.