Macrosolen bidoupensis (Loranthaceae), a new species from Bidoup Nui Ba National Park, southern Vietnam
Author
Tagane, Shuichiro
Center for Asian Conservation Ecology, Kyushu University, 744 Motooka, Fukuoka, 819 - 0395, Japan
stagane29@gmail.com
Author
Dang, Van Son
Author
Ngoc, Nguyen Van
https://orcid.org/0000-0002-1518-9470
Center for Asian Conservation Ecology, Kyushu University, 744 Motooka, Fukuoka, 819 - 0395, Japan & Department of Biology, Dalat University, 01 - Phu Dong Thien Vuong, Dalat, Vietnam
Author
Binh, Hoang Thi
Center for Asian Conservation Ecology, Kyushu University, 744 Motooka, Fukuoka, 819 - 0395, Japan & Department of Biology, Dalat University, 01 - Phu Dong Thien Vuong, Dalat, Vietnam
Author
Komada, Natsuki
Author
Wai, Jarearnsak Sae
Prince of Songkla University, Hat Yai, Thailand & Institute of Ecology and Evolutionary Biology, National Taiwan University, Taipei, Taiwan
Author
Naiki, Akiyo
Iriomote Station, Tropical Biosphere Research Center, University of the Ryukyus, 870 Uehara, Taketomi-cho, Yaeyama-gun, Okinawa, 907 - 1541, Japan
Author
Nagamasu, Hidetoshi
The Kyoto University Museum, Kyoto University, Yoshida Honmachi, Sakyo-ku, Kyoto, 606 - 8501, Japan
Author
Toyama, Hironori
Center for Asian Conservation Ecology, Kyushu University, 744 Motooka, Fukuoka, 819 - 0395, Japan
Author
Yahara, Tetsukazu
Center for Asian Conservation Ecology, Kyushu University, 744 Motooka, Fukuoka, 819 - 0395, Japan
text
PhytoKeys
2017
2017-06-05
80
113
120
http://dx.doi.org/10.3897/phytokeys.80.13338
journal article
http://dx.doi.org/10.3897/phytokeys.80.13338
1314-2003-80-113
FFB1FF87FF94B20BFFB4FFB4FFAAC35C
816502
Macrosolen bidoupensis Tagane & V.S.Dang
sp. nov.
Figures 1
, 2
Diagnosis.
Macrosolen bidoupensis
is similar to
Macrosolen tricolor
Danser of China, Laos and Vietnam in leaf shape and size and by possession of a 2-flowered umbel, but differs in having a sessile to very short petiole (up to 0.7 mm long in
M. bidoupensis
vs. 2-3 mm long in
M. tricolor
), more lateral veins (4 or 5 vs. 2 or 3 pairs), slightly cordate to rounded leaf bases (vs. cuneate) and a basally green corolla tube (vs. reddish around the basal 1/3). The leaf shape is also somewhat similar to
Macrosolen platyphyllus
Danser of Thailand, Peninsular Malaysia and Borneo, but easily distinguished in having much smaller leaves and flowers, and very short petioles.
Figure 1.
Macrosolen bidoupensis
Tagane & V.S.Dang.
A
Branch with flower buds
B
2-flowered inflorescence
C
Flower with corolla tube opened
D
Fresh fruit
E
Dried fruit
F
Seed
G
seed (left) and longitudinal section of seed (right). Materials:
Tagane et al. V4083
(KYO). Drawn by N. Komada.
Figure 2.
Macrosolen bidoupensis
Tagane & V.S.Dang.
A
Branch with flower buds
B
Flower
C
Fruits. Photos by
Tagane et al. V4083
.
Type
.
VIETNAM
.
Lam Dong Province
,
Bidoup Nui Ba National Park
, in lower montane evergreen forest,
12°10'34.7"N
,
108°41'08.4"E
, alt.
1533 m
,
21 Feb. 2016
, with flowers,
Tagane S.
,
Nagamasu H.
,
Naiki A.
,
Dang V. Son
,
Ngyuen V. Ngoc
,
Binh T. Hoang
&
Wai J. V
4083
(holotype-KYO!, fl. spirit collection; isotypes NTU!, the herbarium of
Bidoup Nui Ba National Park
!, VNM!)
.
Description.
Hemi-parasitic shrub, 25-40 cm tall, totally glabrous. Branches terete, grayish to grayish brown, lenticellate. Leaves opposite; blade elliptic, broadly elliptic, broadly ovate-elliptic, or circular, 1.2-4.8
x
1.2-3.5 cm, coriaceous, apex broadly obtuse to rounded, base slightly cordate to rounded, margin entire, grayish green and slightly lustrous adaxially, dull olive abaxially, midrib prominent at basal 1/3 to 1/2 abaxially, secondary veins 4 or 5 pairs, obscure or visible adaxially, obscure to faintly visible abaxially; petiole to 0.7 mm long. Inflorescences axillary, sometimes at older leafless nodes, 2-flowered umbels; peduncle ca. 0.9 mm long. Pedicel 1.1-1.5 mm long; central bract suborbicular, ca. 1 mm long, bracteoles suborbicular, connate, ca. 1 mm long; mature buds 2.8-3.2 cm long; calyx limb (calyculus) annular, ca. 0.7 mm long; corolla green tinged with red at top of tube and lobes, symmetrically 6-winged at about two thirds the length, tube 2.2-2.4 cm long in anthesis, gradually dilated, inflated, slightly curved, lobes lanceolate, 6-8 mm long, reflexed in anthesis; stamens 6, anthers ca. 2.3 mm long, filaments basally adnate to corolla tube, free part 2.3-3.3 mm long; ovary 2.8-3.3 mm long; style 2.8-3.2 cm long, stigma capitate. Berry reddish orange, subglobose to slightly depressed-barrel-shaped, ca. 8 mm high, ca. 6 mm wide, ovoid when dry, apex beaked by calyx limb. Seed 1, ellipsoidal, ca. 5 mm high, ca. 3 mm wide, longitudinally 6-grooved. The measurements of the flower characters are derived from the spirit collection.
Other specimen examined in Vietnam.
Bidoup Nui Ba National Park; in lower montane evergreen forest,
12°10'34.9"N
,
108°41'04.4"E
, alt. 1533 m, 22 Feb. 2016, with fruits,
Tagane et al.V4169
(FU, NTU, the herbarium of Bidoup Nui Ba National Park, VNM).
Phenology.
Flowering and fruiting specimens were collected in February.
Distribution and habitat.
Vietnam (so far known only from Bidoup Nui Ba National Park, Lam Dong Province). In lower montane evergreen forest, ca. 1500 m altitude.
GenBank accession no.
Tagane et al. V4083
: LC259010 (
rbcL
), LC259011 (
matK
). The partial
rbcL
sequence of
M. bidoupensis
was identical to
M. tricolor
(GenBank accession no. HQ317771) of the total 517 bp and differed at 3 bases of the 518 total from
M. cochinchinensis
(KP094775 and HQ317768). The partial
matK
sequence of
M. bidoupensis
differed at 23 bases from
M. cochinchinensis
(EU544439) of the 813 total.
Etymology.
The specific epithet
bidoupensis
is derived from its type locality.
Conservation status.
Least Concern (LC). Bidoup Nui Ba National Park is located on Central highland of Vietnam. It covers ca. 700.38 km2 and is the core zone of the Langbiang biosphere reserve (
Bidoup-Nui Ba National Park 2016
).
Macrosolen bidoupensis
is commonly found in the lower montane evergreen forest in the national park. Since the forest is widely seen and well-protected, we suggest this status as LC according to
IUCN (2012)
.
Note.
A similar species,
Macrosolen tricolor
is known from lower elevations (below 100 m,
Qiu and Gilbert 2003
), and the two species do not occur sympatrically.