An inordinate fondness for Osedax (Siboglinidae: Annelida): Fourteen new species of bone worms from California
Author
Rouse, Greg W.
Author
Goffredi, Shana K.
Author
Johnson, Shannon B.
Author
Vrijenhoek, Robert C.
text
Zootaxa
2018
2018-02-05
4377
4
451
489
journal article
30843
10.11646/zootaxa.4377.4.1
90327333-dbf7-4a3e-9a7e-7c2fd259d963
1175-5326
1165874
C36D839B-A704-41A8-AC2C-2A75AE39F23C
Osedax docricketts
n. sp.
Figure 4E
‘nude-palp-C’ (
Higgs
et al.
2014
;
Rouse
et al.
2011
;
Rouse
et al.
2015
;
Vrijenhoek
et al.
2009
) ‘Sagami-6’ (GenBank
COI
sequence series
FM998088
-
FM998107
; Pradillon et al. unpublished)
Holotype
:
SIO-BIC
A
1644
,
Female
(GenBank
COI
sequence
FJ347626
), fixed in formalin preserved in ethanol, collected from
whale fall deployed
at
1018 meters depth
in
Monterey Submarine Canyon
,
California
(
36°46.308’N
;
122° 4.981’W
),
ROV
Tiburon
dive number
1049,
Oct. 25
, 2006.
Diagnosis and description.
Holotype incomplete female; preserved trunk 0. 5 mm long, 0.2 mm wide; crown of four palps, 2 mm long (
Fig. 4E
). Tube gelatinous cylinder around trunk and base of palps. Oviduct not discerned. In life, palps white with some reddish patches. Palps without obvious pinnules (
Fig. 4E
). Trunk with no obvious demarcation into upper and lower trunk, or any pigmentation. Ovisac mostly missing in holotype, some traces of roots present. Males not seen.
Distribution.
Known from Monterey Bay, California from 1820 meters depth, and Sagami Bay, Japan (
Table 2
). It has been
found living on cow and whale bones.
Etymology.
This species is named (noun in apposition) for the ROV
Doc Ricketts
, which was used to collect many
Osedax
specimens.
Remarks.
Osedax docricketts
n. sp.
is part of
Osedax
Clade II, which all show apinnulate palps, and is closest relative to the clade comprised of
Osedax westernflye
r
n.
sp.
and
O. knutei
n. sp.
(minimum divergence of 15%) though it shows smaller distance (14%) to other taxa such as
O. lonnyi
n. sp.
(
Table 4
). The four available
COI
sequences for
Osedax docricketts
n. sp.
from California (
Table 3
) show less that 1% sequence divergence. The only retained specimen is the holotype as the others were destroyed for sequencing and the holotype itself is only a partial specimen. Twenty additional
COI
sequences were referred to
Osedax
sp. Sagami-6 (Pradillon et al., GenBank, unpublished). Together, the four California and 11 of the Japanese sequences (
FM998088
,
FM998089
,
FM998091
,
FM998093
,
FM998094
,
FM998098
,
FM998099
,
FM998100
,
FM998101
,
FM998103
,
FM998107
) encompass less than 1% sequence divergence; so, we propose that
Osedax docricketts
n. sp.
is also found in Japan. The nine remaining Japanese specimens with>2.4% uncorrected pairwise distance from the 11 Japanese sequences (GenBank sequences:
FM998090
,
FM998092
,
FM998095
,
FM998096
,
FM998097
,
FM998102
,
FM998104
,
FM998105
,
FM998106
) require further investigation before their assignment to
Osedax docricketts
n. sp.
can be justified. See
Table 4
for comparison of California
Osedax docricketts
n. sp.
versus Japanese Sagami-6 representative sequences. There is little to distinguish
Osedax docricketts
n. sp.
from other Clade II species.