Spiky pygmy devils: revision of the genus Discotettix (Orthoptera: Tetrigidae) and synonymy of Discotettiginae with Scelimeninae Author Skejo, Josip Author Pushkar, Taras I. Author Kasalo, Niko Author Pavlović, Marko Author Deranja, Maks Author Adžić, Karmela Author Tan, Ming Kai Author Rebrina, Fran Author Muhammad, Amira Aqilah Author Abdullah, Nurul Ashikin Author Japir, Razy Author Chung, Arthur Y. C. Author Tumbrinck, Josef text Zootaxa 2022 2022-12-06 5217 1 1 64 http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN journal article 203364 10.11646/zootaxa.5217.1.1 6b4363b1-e59c-49b2-b32d-3928329d8679 1175-5326 7403418 86CD1EDF-8C38-4A90-888A-185B8481A6ED Subgenus Mnesarchus Stål, 1877 , nom. resurr. Mnesarchus Stål, 1877: 55 ; synonymized with Discotettix by Bolívar (1887) . Mnesarchus (as subgenus of Discotettix ): Kevan 1966: 380 . Discotettix (partim): Bolívar 1887: 306 ; Kirby 1910: 2 [listed in the catalog]; Blackith 1992: 46 [listed in the catalog]; Yin et al. 1996: 866 [listed in the catalog], Otte 1997: 32 [listed in the catalog]. Type species: Mnesarchus scabridus Stål, 1877 , by original monotypy. Taxonomic notes. The genus Mnesarchus have been synonymized with Discotettix by Bolívar (1887) . Kevan (1966) recognized it as distinct subgenus but later Mnesarchus has been considered a synonym of Discotettix again ( Blackith, 1992 ; Yin et al. , 1996 ; Otte, 1997 ). The type species is morphologically very different from other Discotettix members and is thus assigned to its own subgenus, Mnesarchus . It can be easily separated from other species of the genus by the set of the following characters: (I) frontal costa bifurcates at the lower margin of the compound eyes (bifurcation is in the lower third of the compound eyes in D. aruanus sp. n. , D. belzebuth , D. doriae , D. kirscheyi sp. n. , D. selysi , and D. sumatrensis sp. n. ), (II) FM present as a low tubercle (developed and elevated in D. aruanus sp. n. , D. belzebuth , D. doriae , D. kirscheyi sp. n. , D. selysi , and D. sumatrensis sp. n. ), (III) MM laterally compressed and elevated (spine-like in D. belzebuth , triangular protrusions in D. sumatrensis sp. n. , D. kirscheyi sp. n. , while very low in D. aruanus sp. n. , D. doriae and D. selysi ), (IV) MML low and triangular, compressed elevations (spine-like in D. belzebuth , triangular protrusions in D. sumatrensis sp. n. , similarly formed in D. selysi ), (V) interscapular area triangular, with large concavity (similarly to D. selysi , and D. sumatrensis sp. n. , almost parallel in D. belzebuth ), (VI) lateral and humeral carinae strongly toothed (similar to D. belzebuth , granulated in D. selysi , and D. sumatrensis sp. n. ), (VII) weak ML (almost absent in D. scabridus , while well visible in other species), (VIII) VL complex, with a few spines (in other species usually with one main spine, and a saw-like margin) and (IX) tegmen more elongated ( TL/TW > 5) than in the other species of the genus ( TL/TW <3.5). Composition. The type species of Mnesarchus is the only species presently assigned to this subgenus.