Revision of the Megasoma (Megasoma) gyas (Jablonsky in Herbst, 1785) species group (Coleoptera, Scarabaeidae, Dynastinae) Author Prandi, Massimo Via Del Seminario 16, I- 25087, Salo, BS, Italy https://orcid.org/0000-0002-4671-3589 prandi.m@libero.it Author Grossi, Paschoal C. Universidade Federal Rural de Pernambuco, Departamento de Agronomia / Fitossanidade, Laboratorio de Taxonomia de Insetos, Programa de Pos-graduacao em Entomologia Agricola-PPGEA, 52171 - 900, Recife, Pernambuco, Brazil Author Vaz-De-Mello, Fernando Z. Universidade Federal de Mato Grosso, Instituto de Biociencias, Departamento de Biologia e Zoologia, Colecao Entomologica, 78060 - 900, Cuiaba, Mato Grosso, Brazil text ZooKeys 2020 999 109 145 http://dx.doi.org/10.3897/zookeys.999.53130 journal article http://dx.doi.org/10.3897/zookeys.999.53130 1313-2970-999-109 B8D6EF7CBFED4758A1EBA58F1711BD30 833665678A4F5EEF87330B7783687C86 Megasoma (Lycophontes) joergenseni penyai Nagai, 2003 Figure 27 Megasoma joergenseni ssp. penyai Nagai, 2003: 38-39. Remarks. The subspecies penyai is restricted to an area in central-western Paraguay (holotype from Loma Plata, Chaco). The main characters differentiating it from the subspecies joergenseni are found in the smaller median thoracic horn and in the denser pubescence, giving a more brownish color. It is also usually smaller than ssp. joergenseni . (Fig. 27 , courtesy of Mushi-sha). Figure 27. A M. joergenseni peynai ♂ B M. joergenseni peynai ♀.