Pyrgotid Flies Assigned To Apyrgota. Iii. Species Of Afropyrgota Gen. N. And Tylotrypes (Diptera, Pyrgotidae) Author Korneyev, V. A. text Vestnik Zoologii 2015 2015-02-01 49 1 25 40 https://www.degruyter.com/doi/10.1515/vzoo-2015-0003 journal article 10.1515/vzoo-2015-0003 642f8951-9cc2-4a64-b928-9cb210dd9a6a 2073-2333 6404027 Afropyrgota V. Korneyev , gen. n. Type species: Apyrgota marshalli Hendel, 1914 . D i a g n o s i s. Robust, wasp-like flies, very similar to Eupyrgota in having large subocular sclerite, bare scutellum, wing with spurious vein inserted to deep flexion of R 2+3 , femora with two ventral rows of shortened and thickened setae on ventral side; mostly bare epandrium with elongated surstyli (as in Eupyrgota s. str. ), oviscape with opening directed ventrally, paired rows of stiff setulae located anterolaterally of it, and short, T- or Y-shaped aculeus with strongly widened shoulders basally. They differ from Eupyrgota by the saddlelike prosternum without nipple or finger-like projections ventrally, and oviscape bearing no postero-laterally directed sclerotized hooks. D e s c r i p t i o n. Afrotropical Region (from Nigeria to Kenya and Tanzania ). Head. Frons and facialium broad, wider than eye, face with narrow, often very low carina; epistome low; subocular sclerite large; setae moderately short: oc, or and vte shorter than or at most as long as poc. Thorax. Presternum saddle-like, without nipple-like ventral projections. Scutellum bare, with two pairs of setae. Legs. Female fore and mid trochanters without thickened setulae; mid femur of female with or without femoral organ; setae of apicoventral rows moderately thickened and shortened. Wing usually grey microtrichose, sometimes widely or almost entirely yellow tinged; subcostal vein straight, interrupted at fusion with costa. Pterostigma short. Vein R 2+3 deeply and sharply bent at insertion of spurious vein, R 4+5 bare. Costal vein reaching apex of M. Cell bcu with calypters with fine and short fringe, without spinulose cilia. Abdomen. Sternites 1 and 2 entirely fused, forming long Y-shaped synsternite 1+2. Male genitalia. Hypandrium with phallic guide. Epandrium almost entirely bare, with elongate lateral surstyli. Medial surstyli without prensisetae. Female genitalia. Oviscape shorter than abdomen; its opening ventrally directed; apical fields of trichoid sensilla present: on lateroventral lobe (with thick setulae arranged in two rows — fig. 2, 2 : lvl; 4, 9; 8, 4 ) and dorsomedial lobe ( fig. 2, 2 : dl ); medioventral part at the apex widely desclerotized ( fig. 2 , 1 ; 4 , 8; 8 , 4 ). Sclerotized hooks distal of them absent. Aculeus short, strongly widened in basal one-third and narrowly stiletto-like in apical portion (Y-like, see fig. 2 , 3 ; 4 , 10 ; 8 , 5 ). Ventral receptacle tubular, with spherical apical portion ( fig. 2 , 4 ). One or two unequal smooth, poorly sclerotized, slightly curved spermathecae observed in few dissected specimens (related species of Eupyrgota usually have 2 larger and one smaller spermatheca, and I suppose this could be actual number of spermathecae in Afropyrgota , too, while the other could be lost during preparation). Key to species of Afropyrgota 1. Orbit with opalescent or matt, microtrichose area at level of antenna base ( fig. 7 , 2 ). Wing yellow with slightly darker apex ( fig 7 , 1 , 5 ). Thorax, femora and abdomen dark reddish brown, black setulose. Scutellum with 3 pairs of sctl ( fig. 7 , 4 ). WL> 15 mm . Tanzania . ....................... Afropyrgota uzungwa sp. n. — Orbit entirely shining. WL < 15 mm . Other characters variable. ............................................................... 2. 2. Mesonotum densely and long black setulose ( fig. 3 , 2 ). Scutellum with 3 pairs of sctl ( fig. 3 , 5 ). Proepisternal ridge with 8–10 long setae. Oviscape with 2 yellow hook-like, anteromesally directed lobes ( fig. 3 , 9 ). Wing grey microtrichose with two darker marks. Female mid femur yellowish setulose, without femoral organ ( fig. 3 , 7 ). Nigeria . ............................................................... Afropyrgota mambilla sp. n. — Mesonotum moderately short reddish brown setulose. Scutellum with 1–2 pairs of sctl. Proepisternal ridge with 2–6 moderately short setae. Oviscape without such lobes. Wing and mid femur variable. ..... 3. 3. Scutellum with one pair of sctl ( fig. 1 , 5 ). Frons widely black. Mid femur with very large cuneiform femoral organ almost entirely covering its apical half ( fig. 1 , 7 ). Kenya . ..... Afropyrgota copelandi sp. n. — Scutellum with two pairs of sctl. Frons and mid femur variable. ............................................................... 4. 4. Body entirely reddish yellow setulose, including pleura, abdomen and legs. Mid femur with cuneiform femoral organ narrower than femur ( fig. 6, 6 ). Nigeria , Kenya . .............................. Afropyrgota ota sp. n. — Setae on pleura, abdomen and legs black or dark brown. Mid femur without femoral organ, densely brown setulose ( fig. 4 , 7 ). Nigeria , Kenya , DR Congo ( Zaïre ). ............... Afropyrgota marshalli (Hendel)