A review of the genus Pupopsis Gredler, 1898 (Gastropoda: Stylommatophora: Enidae), with the descriptions of eight new species from China
Author
Wu, Min
Author
Gao, Linhui
text
Zootaxa
2010
2725
1
27
journal article
46921
10.5281/zenodo.199982
558b2fa1-879c-44e4-956d-9abd9c9a249a
1175-5326
199982
6.
Pupopsis torquilla
(
Möllendorff, 1901
)
Figs. 1
,
3
C–D, 8A–8B;
Table 2
Buliminus
(
Pupopsis
)
torquilla
Möllendorff, 1901
: 376
, pl. 17, figs. 19, 20.
Buliminus
(
Pupopsis
)
torquilla—
Kobelt, 1902: 876, pl. 124, figs. 19, 20.
Pupopsis
torquilla—
Yen, 1939
: 90
, pl. 8, fig. 41.
Type
material.
Lectotype
: SMF-42082, Tshiu-dsei-dsy near Hsi-ku-tsheng, Kansu (today's Zhouqu County, 33°48ʹN, 104°18ʹE, Gansu),
China
; Potanin 206, Mlldff Colln. (measurements of
lectotype
: ewh 1.500, wh 8.625, h 8.1, diam. maj. 4.2, aph 3.0, apw 2.7). Whereabouts of additional
type
material unknown.
Type
locality.
Tshiu-dsei-dsy near Hsi-gu-tshêng (Gansu).
Material examined.
HBUMM-05584: 2 fms,
829 m
a.s.l., 32°51ʹ44.7ʹ N, 104°50ʹ48.2ʹ E,
Hendan
Village, on a nearby hill located on the opposite bank of Baishuijiang River and national highway o. 212, S. Gansu,
China
,
2006–IX–29
, leg. M. Wu, J. Liu, W. Zheng & L. Gao.
Distribution.
S. Gansu.
Diagnosis.
Shell with 7.500–8.625 whorls. Height 7.3–8.1, diam. maj. 3.5–4.2. Parietal tooth present. Columellar margin with one blunt tooth. Palatal wall with a distinct or indistinct ridge-like tooth.
Re-description of shell.
Cylindrical-conic; apex not acuminate or slightly but distinctly acuminate (in
holotype
); dextral; thin-shelled; solid; opaque; glossy; with 7.500–7.750–8.000 whorls. Whorls spirally grooved (densely and evenly spaced near umbilical region, in
lectotype
) or not (in newly collected specimens); convex.
Embryonic shell smooth; not polished; with 1.375–1.438–1.500 whorls. Postnuclear whorls wrinkled. Suture simple. Last whorl straight in front; rounded at periphery. Shell 7.3–7.4–7.5 high; 3.5–3.6–
3.7 in
diam. maj. Height/diam. maj. ratio 1.99–2.07–2.14. Aperture semicircular; oblique; its insertions separated; 2.8–2.8–2.8 high, 2.4–2.4–2.5 wide. Ratio of shell height to aperture height 2.62–2.65–2.68. Palatal tooth present, ridge-like, strongly (in
lectotype
) or indistinctly (in newly collected specimens) developed. If distinct, palatal ridge-like tooth entering spirally approximately 0.250 whorl. Palatal margin rounded. Peristome expanded; thickened. Parietal callus indistinct. Angular tubercle present. Parietal tooth much larger than angular tubercle, extending inwards. Columellar margin reflexed; with one blunt tooth which is expanded inwards. Umbilicus more or less widely open. Shell, including upper whorls, uniformly corneous-brown; aperture white.
Remarks.
Möllendorff (1901)
doesn’t mention a
holotype
or any other kind of
types
in his original description of
Buliminus
(
Pupopsis
)
torquilla
, nor does he indicate how many specimens belonged to the
type
series.
Yen (1939: 90)
refers to the only specimen of
B. torquilla
in colln. Möllendorff in SMF (SMF 42082) as the
holotype
of the taxon. Since there is no evidence that this specimen constitutes the entire
type
series, it is not the
holotype
but became, by Yen referring to it as the
holotype
, the
lectotype
of
B. torquilla
Möllendorff, 1901
(ICZN Art 74.6).
Our newly collected material differs from the original description of
P
.
torquilla
in the weakness or absence of apertural teeth and in having fewer whorls.