A mountain of millipedes VII: The genus Eviulisoma Silvestri, 1910, in the Udzungwa Mountains, Tanzania, and related species from other Eastern Arc Mountains. With notes on Eoseviulisoma Brolemann, 1920, and Suohelisoma Hoffman, 1963 (Diplopoda, Polydesmida, Paradoxosomatidae) Author Enghoff, Henrik text European Journal of Taxonomy 2018 2018-06-19 445 1 90 journal article 10.5852/ejt.2018.445 5dfd4ff4-6c8b-47f4-9aa7-0f6b1b2af967 1489598 852A3F68-B728-413A-B12E-56F306D56C35 The kwabuniense group Diagnosis Species of Eviulisoma in which male sternum 6 is deeply excavated with unlobed margins, there is an intermediate acropodital process and the solenophore is a relatively open sheet with mostly three, more rarely two apical lobes. Fig. 8. Gonopods in Eviulisoma spp. A . E. akkariae sp. nov. , paratype, right gonopod, colour-coded: light green = coxa; dark green = cannula; light blue = prefemoral part; purplish blue = mesal acropodital process; pink = intermediate acropodital process; orange = solenophore; yellow = solenomere. B . E. culter sp. nov. , paratype, right gonopod, solenomere entering solenophore. C . E. ottokrausi sp. nov. , paratype, left gonopod, solenomere dislodged from solenophore. Scale bars: A = 0.1 mm; B–C = 0.05 mm. Table 3. Species groups of Eviulisoma Silvestri, 1910 , their constituent species and main morphological characters. Note that with the exception of E. alluaudi Brolemann, 1920 , E. jeanneli Brolemann, 1920 and E. silvaticum Attems, 1953 , which are tentatively assigned to the kwabuniense group, the table only includes species from the Udzungwa, Rubeho and Kanga Mts.
CHARACTERS
sternum 6 margins of sternum acropodite with intermediate mesal solenophore
excavation and lobe 6 excavation a separate basal acropodital acropodital
Group name Included species between coxae 4 ‘femorite’ process process
kwabuniense group E. kwabuniense + & + not lobed no + not compressed large, not rolled, with
E. acaciae sp. nov. (two-)three apical lobes
E. aequilobatum sp. nov.
E. akkariae sp. nov.
E. cetafi sp. nov.
E. chitense sp. nov.
E. commelina sp. nov.
E. ejti sp. nov.
E. kalimbasiense sp. nov.
E. navuncus sp. nov.
E. nessiteras sp. nov.
E. ottokrausi sp. nov.
E. paradisiacum sp. nov.
( E. alluaudi )
( E. jeanneli )
( E. silvaticum )
dabagaense group E. dabagaense + & + not lobed no strongly laterally tubular, ‘rolled sheet’
E. grumslingslak sp. nov. compressed
E. coxale sp. nov.
E. culter sp. nov.
sternale group E. sternale sp. nov. + & + lobed no not compressed tubular, ‘rolled sheet’
E. zebra sp. nov.
E. kangaense sp. nov.
iuloideum group E. iuloideum + & + not lobed yes not compressed large, not rolled, no
E. articulatum sp. nov. apical lobes
Ungrouped E. angulatum sp. nov. + & + angled no not compressed large, not rolled, two apical lobes
E. breviscutum sp. nov. - & - NA no not compressed, short, shield-like large, not rolled, two apical lobes
E. biquintum sp. nov. - & - NA no not compressed large, rolled
The included species furthermore share a hypoproct with three prominent marginal tubercles. Most of the species are apparently pallid (although fading of colour cannot be excluded), only E. akkariae sp. nov. , which is also larger than the others, having a distinct ringed colour pattern. Included species:
E. kwabuniense Kraus, 1958 E. acaciae sp. nov. E. aequilobatum sp. nov. E. akkariae sp. nov. E. cetafi sp. nov. E. chitense sp. nov. E. commelina sp. nov. E. ejti sp. nov. E. kalimbasiense sp. nov. E. navuncus sp. nov. E. nessiteras sp. nov. E. ottokrausi sp. nov. E. paradisiacum sp. nov. E. alluaudi Brolemann, 1920 , E. jeanneli Brolemann, 1920 , and E. silvaticum Attems, 1953 , may also be tentatively included in this group, based on published descriptions ( Attems 1953 ; Brolemann 1920 ; Jeekel 2003 ; VandenSpiegel & Golovatch 2014 ).