<document ID-DOI="10.11646/zootaxa.5250.1.1" ID-ISSN="1175­5334" ID-Zenodo-Dep="7705657" ID-ZooBank="4729EAF3-8B78-4AAD-B373-9179D008F239" approvalRequired="130" approvalRequired_for_document="1" approvalRequired_for_originalDoi="1" approvalRequired_for_textStreams="123" approvalRequired_for_treatments="5" checkinTime="1678178160677" checkinUser="plazi" docAuthor="WANG, YA-LIAN, LI, BING-XUE, CUI, MENG-DI, ZHANG, YUAN, WANG, LE, ZHANG, CAI-HONG, TSAUR, SHUN-CHERN, CHEN, HONG-WEI &amp; HUANG, JIA" docDate="2023" docId="0393914C2A10874AC282FF1E6A63F9C3" docLanguage="en" docName="zootaxa.5250.1.1.pdf" docOrigin="Zootaxa 5250 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5250.1.1" docStyle="DocumentStyle:8B0D3ECF822058C8413568C103B59429.5:Zootaxa.2001-2006.monograph" docStyleId="8B0D3ECF822058C8413568C103B59429" docStyleName="Zootaxa.2001-2006.monograph" docStyleVersion="5" docTitle="Stegana (Steganina) hirticeps Wang, Gao, &amp; Chen 2013" docType="treatment" docVersion="3" lastPageNumber="51" masterDocId="FFAAE9342A228778C215FF896958FB43" masterDocTitle="Revision of the subgenus Stegana (Steganina) from China, with assessment of species delimitation using DNA barcodes (Diptera, Drosophilidae)" masterLastPageNumber="109" masterPageNumber="1" pageNumber="51" updateTime="1678205790159" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title>Revision of the subgenus Stegana (Steganina) from China, with assessment of species delimitation using DNA barcodes (Diptera, Drosophilidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>WANG, YA-LIAN</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>LI, BING-XUE</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>CUI, MENG-DI</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>ZHANG, YUAN</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>WANG, LE</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>ZHANG, CAI-HONG</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>TSAUR, SHUN-CHERN</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>CHEN, HONG-WEI</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>HUANG, JIA</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>Zootaxa</mods:title>
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<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-03-07</mods:number>
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<mods:detail type="volume">
<mods:number>5250</mods:number>
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<mods:number>1</mods:number>
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<mods:url>http://dx.doi.org/10.11646/zootaxa.5250.1.1</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.11646/zootaxa.5250.1.1</mods:identifier>
<mods:identifier type="ISSN">1175­5334</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.7705557" ID-Zenodo-Dep="7705557" LSID="urn:lsid:plazi:treatment:0393914C2A10874AC282FF1E6A63F9C3" httpUri="http://treatment.plazi.org/id/0393914C2A10874AC282FF1E6A63F9C3" lastPageNumber="51" pageId="50" pageNumber="51">
<subSubSection box="[151,844,151,178]" pageId="50" pageNumber="51" type="nomenclature">
<paragraph blockId="50.[151,844,151,213]" box="[151,844,151,178]" pageId="50" pageNumber="51">
<heading bold="true" box="[151,844,151,178]" fontSize="11" level="1" pageId="50" pageNumber="51" reason="1">
<taxonomicName authority="Wang, Gao, &amp; Chen, 2013" authorityName="Wang, Gao, &amp; Chen" authorityYear="2013" box="[151,844,151,178]" class="Insecta" family="Drosophilidae" genus="Stegana" kingdom="Animalia" order="Diptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="hirticeps" subGenus="Steganina">
<emphasis bold="true" box="[151,844,151,178]" pageId="50" pageNumber="51">
<emphasis bold="true" box="[151,249,151,177]" italics="true" pageId="50" pageNumber="51">Stegana</emphasis>
(
<emphasis bold="true" box="[266,388,151,177]" italics="true" pageId="50" pageNumber="51">Steganina</emphasis>
) 
<emphasis bold="true" box="[405,509,151,178]" italics="true" pageId="50" pageNumber="51">hirticeps</emphasis>
Wang, Gao, &amp; Chen, 2013
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[151,249,187,213]" pageId="50" pageNumber="51" type="description">
<paragraph blockId="50.[151,844,151,213]" box="[151,249,187,213]" pageId="50" pageNumber="51">
(
<figureCitation box="[159,241,187,213]" captionStart="FIGURE 23" captionStartId="50.[151,250,1506,1530]" captionTargetBox="[159,1429,673,1474]" captionTargetId="figure-167@50.[151,1436,664,1483]" captionTargetPageId="50" captionText="FIGURE 23. Stegana (Steganina) hirticeps Wang, Gao &amp;, Chen, 2013 from Nanling, Shaoguan, Guangdong, China, male terminalia. A. Epandrium, surstylus, and cercus in lateral view; B. C. Hypandrium, pregonite, aedeagus, and phallapodeme in ventral and lateral views; D, E. Aedeagal sheath in ventral and lateral views; F. Subepandrial sclerite in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7705721" httpUri="https://zenodo.org/record/7705721/files/figure.png" pageId="50" pageNumber="51">Fig. 23</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection box="[151,939,256,280]" pageId="50" pageNumber="51" type="reference_group">
<paragraph blockId="50.[151,939,256,280]" box="[151,939,256,280]" pageId="50" pageNumber="51">
<treatmentCitationGroup box="[151,939,256,280]" pageId="50" pageNumber="51">
<taxonomicName authority="Wang, Gao, &amp; Chen" authorityName="Wang, Gao, &amp; Chen" authorityYear="2013" box="[151,663,256,280]" class="Insecta" family="Drosophilidae" genus="Stegana" kingdom="Animalia" order="Diptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="hirticeps" subGenus="Steganina">
<emphasis box="[151,233,256,279]" italics="true" pageId="50" pageNumber="51">Stegana</emphasis>
(
<emphasis box="[248,349,256,279]" italics="true" pageId="50" pageNumber="51">Steganina</emphasis>
) 
<emphasis box="[364,453,256,280]" italics="true" pageId="50" pageNumber="51">hirticeps</emphasis>
Wang, Gao, &amp; Chen
</taxonomicName>
in 
<treatmentCitation author="Wang, L. &amp; Gao, J. J. &amp; Chen, H. W." box="[696,935,256,280]" httpUri="http://treatment.plazi.org/id/038EBF77FFD81874FE09976EC195FA57" page="2002" pageId="50" pageNumber="51" year="2013">
<bibRefCitation author="Wang, L. &amp; Gao, J. J. &amp; Chen, H. W." box="[696,935,256,280]" pageId="50" pageNumber="51" pagination="1993 - 2013" refId="ref74574" refString="Wang, L., Gao, J. J. &amp; Chen, H. W. (2013) Ten new species of the subgenus Steganina (Diptera: Drosophilidae: Stegana) from southern China. Journal of Natural History, 47, 1993 - 2013. https: // doi. org / 10.1080 / 00222933.2012.763055" type="journal article" year="2013">
Wang 
<emphasis box="[760,812,256,280]" italics="true" pageId="50" pageNumber="51">et al.</emphasis>
2013: 2002
</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection pageId="50" pageNumber="51" type="discussion">
<paragraph blockId="50.[151,1437,326,640]" pageId="50" pageNumber="51">
<emphasis bold="true" box="[151,267,326,352]" pageId="50" pageNumber="51">Remarks.</emphasis>
The specimens collected from 
<collectingRegion box="[639,770,326,352]" country="China" name="Guangdong" pageId="50" pageNumber="51">Guangdong</collectingRegion>
and 
<collectingRegion box="[832,918,326,352]" country="China" name="Yunnan" pageId="50" pageNumber="51">Yunnan</collectingRegion>
slightly differ from the 
<typeStatus box="[1208,1256,326,352]" pageId="50" pageNumber="51">type</typeStatus>
materials from 
<collectingRegion box="[151,247,362,388]" country="China" name="Guangxi" pageId="50" pageNumber="51">Guangxi</collectingRegion>
in having the aedeagus with several minute, serrated processes laterally (
<figureCitation box="[1065,1199,362,388]" captionStart="FIGURE 23" captionStartId="50.[151,250,1506,1530]" captionTargetBox="[159,1429,673,1474]" captionTargetId="figure-167@50.[151,1436,664,1483]" captionTargetPageId="50" captionText="FIGURE 23. Stegana (Steganina) hirticeps Wang, Gao &amp;, Chen, 2013 from Nanling, Shaoguan, Guangdong, China, male terminalia. A. Epandrium, surstylus, and cercus in lateral view; B. C. Hypandrium, pregonite, aedeagus, and phallapodeme in ventral and lateral views; D, E. Aedeagal sheath in ventral and lateral views; F. Subepandrial sclerite in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7705721" httpUri="https://zenodo.org/record/7705721/files/figure.png" pageId="50" pageNumber="51">Fig. 23B, C</figureCitation>
); the aedeagal sheath lacking serrated process (
<figureCitation box="[436,571,398,424]" captionStart="FIGURE 23" captionStartId="50.[151,250,1506,1530]" captionTargetBox="[159,1429,673,1474]" captionTargetId="figure-167@50.[151,1436,664,1483]" captionTargetPageId="50" captionText="FIGURE 23. Stegana (Steganina) hirticeps Wang, Gao &amp;, Chen, 2013 from Nanling, Shaoguan, Guangdong, China, male terminalia. A. Epandrium, surstylus, and cercus in lateral view; B. C. Hypandrium, pregonite, aedeagus, and phallapodeme in ventral and lateral views; D, E. Aedeagal sheath in ventral and lateral views; F. Subepandrial sclerite in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7705721" httpUri="https://zenodo.org/record/7705721/files/figure.png" pageId="50" pageNumber="51">Fig. 23D, E</figureCitation>
). They were conservatively identified as conspecific.
</paragraph>
<paragraph blockId="50.[151,1437,326,640]" pageId="50" pageNumber="51">
<emphasis bold="true" box="[199,460,433,459]" pageId="50" pageNumber="51">Specimens examined.</emphasis>
<materialsCitation collectingDate="2012-05-07" collectingDateMax="2018-05-29" collectingDateMin="2012-05-07" collectionCode="SCAU" collectorName="Y. Lin &amp; H. W. Chen" country="China" county="Nanling" elevation="1300" latitude="21.838057" location="Husa" longLatPrecision="21" longitude="100.09667" municipality="Shaoguan" pageId="50" pageNumber="51" specimenCount="3" specimenCount-female="3" stateProvince="Guangdong">
1♁ (
<collectionCode box="[527,606,434,460]" pageId="50" pageNumber="51">SCAU</collectionCode>
, No. 125523, DNA #2604), 
<collectingCountry box="[950,1043,434,460]" name="China" pageId="50" pageNumber="51">CHINA</collectingCountry>
: 
<collectingCounty box="[1059,1152,434,460]" pageId="50" pageNumber="51">Nanling</collectingCounty>
, 
<collectingMunicipality box="[1169,1282,434,460]" pageId="50" pageNumber="51">Shaoguan</collectingMunicipality>
, 
<collectingRegion box="[1299,1433,434,460]" country="China" name="Guangdong" pageId="50" pageNumber="51">Guangdong</collectingRegion>
, 
<geoCoordinate box="[151,284,470,497]" degrees="24" direction="north" minutes="56" orientation="latitude" pageId="50" pageNumber="51" precision="15" seconds="41" value="24.944721">24°56'41''N</geoCoordinate>
, 
<geoCoordinate box="[294,437,470,496]" degrees="113" direction="east" minutes="03" orientation="longitude" pageId="50" pageNumber="51" precision="15" seconds="04" value="113.05112">113°03'04''E</geoCoordinate>
, alt. 
<quantity box="[488,568,470,496]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="50" pageNumber="51" unit="m" value="1300.0">
<elevation box="[488,568,470,496]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="50" pageNumber="51" unit="m" value="1300.0">1300m</elevation>
</quantity>
, 
<date box="[579,691,470,496]" pageId="50" pageNumber="51" value="2018-05-29">
<collectingDate box="[579,691,470,496]" pageId="50" pageNumber="51" value="2018-05-29">29.v.2018</collectingDate>
</date>
, ex tree trunk, 
<collectorName box="[852,923,470,496]" pageId="50" pageNumber="51">Y. Lin</collectorName>
; 1♁ (
<collectionCode box="[982,1061,470,496]" pageId="50" pageNumber="51">SCAU</collectionCode>
, No. 125524, DNA #2680), 
<location LSID="urn:lsid:plazi:treatment:0393914C2A10874AC282FF1E6A63F9C3:8EE576812A10874AC749FE5F6CC0FAB3" box="[1372,1432,470,496]" country="China" county="Nanling" latitude="21.838057" longLatPrecision="21" longitude="100.09667" municipality="Shaoguan" name="Husa" pageId="50" pageNumber="51" stateProvince="Guangdong">Husa</location>
, 
<location LSID="urn:lsid:plazi:treatment:0393914C2A10874AC282FF1E6A63F9C3:8EE576812A10874AC282FE736840F957" box="[151,280,506,532]" country="China" county="Nanling" latitude="21.838057" longLatPrecision="21" longitude="100.09667" municipality="Shaoguan" name="Longchuan" pageId="50" pageNumber="51" stateProvince="Guangdong">Longchuan</location>
, 
<collectingRegion box="[292,381,506,532]" country="China" name="Yunnan" pageId="50" pageNumber="51">Yunnan</collectingRegion>
, 
<geoCoordinate box="[394,527,506,532]" degrees="24" direction="north" minutes="28" orientation="latitude" pageId="50" pageNumber="51" precision="15" seconds="21" value="24.4725">24°28'21''N</geoCoordinate>
, 
<geoCoordinate box="[541,671,506,533]" degrees="97" direction="east" minutes="54" orientation="longitude" pageId="50" pageNumber="51" precision="15" seconds="06" value="97.901665">97°54'06''E</geoCoordinate>
, alt. 
<quantity box="[729,809,506,532]" metricMagnitude="3" metricUnit="m" metricValue="1.34" pageId="50" pageNumber="51" unit="m" value="1340.0">
<elevation box="[729,809,506,532]" metricMagnitude="3" metricUnit="m" metricValue="1.34" pageId="50" pageNumber="51" unit="m" value="1340.0">1340m</elevation>
</quantity>
, 
<date box="[822,931,506,532]" pageId="50" pageNumber="51" value="2017-11-07">
<collectingDate box="[822,931,506,532]" pageId="50" pageNumber="51" value="2017-11-07">7.xi.2017</collectingDate>
</date>
, ex tree trunk, 
<collectorName box="[1106,1237,506,532]" pageId="50" pageNumber="51">H.W. Chen</collectorName>
; 1♁, 
<specimenCount box="[1301,1336,506,532]" pageId="50" pageNumber="51" type="female">3♀</specimenCount>
(
<collectionCode box="[1353,1432,506,532]" pageId="50" pageNumber="51">SCAU</collectionCode>
, Nos. 125525–28, DNA #2679), Hesong, Menghai, 
<collectingRegion box="[720,809,542,568]" country="China" name="Yunnan" pageId="50" pageNumber="51">Yunnan</collectingRegion>
, 
<geoCoordinate box="[819,952,542,569]" degrees="21" direction="north" minutes="50" orientation="latitude" pageId="50" pageNumber="51" precision="15" seconds="17" value="21.838057">21°50'17''N</geoCoordinate>
, 
<geoCoordinate box="[963,1107,542,569]" degrees="100" direction="east" minutes="05" orientation="longitude" pageId="50" pageNumber="51" precision="15" seconds="48" value="100.09667">100°05'48''E</geoCoordinate>
, alt. 
<quantity box="[1160,1240,542,568]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="50" pageNumber="51" unit="m" value="1600.0">
<elevation box="[1160,1240,542,568]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="50" pageNumber="51" unit="m" value="1600.0">1600m</elevation>
</quantity>
, 
<date box="[1251,1350,542,568]" pageId="50" pageNumber="51" value="2012-05-07">
<collectingDate box="[1251,1350,542,568]" pageId="50" pageNumber="51" value="2012-05-07">7.v.2012</collectingDate>
</date>
, ex tree trunks, 
<collectorName box="[235,363,578,604]" pageId="50" pageNumber="51">H.W. Chen</collectorName>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection box="[199,827,613,640]" pageId="50" pageNumber="51" type="distribution">
<paragraph blockId="50.[151,1437,326,640]" box="[199,827,613,640]" pageId="50" pageNumber="51">
<emphasis bold="true" box="[199,353,613,639]" pageId="50" pageNumber="51">Distribution.</emphasis>
<collectingCountry box="[359,426,614,640]" name="China" pageId="50" pageNumber="51">China</collectingCountry>
(
<collectingRegion box="[441,579,614,640]" country="China" name="Guangdong" pageId="50" pageNumber="51">Guangdong</collectingRegion>
*, 
<collectingRegion box="[601,700,614,640]" country="China" name="Guangxi" pageId="50" pageNumber="51">Guangxi</collectingRegion>
, 
<collectingRegion box="[711,805,614,640]" country="China" name="Yunnan" pageId="50" pageNumber="51">Yunnan</collectingRegion>
*).
</paragraph>
</subSubSection>
</treatment>
</document>