Ceratophila, a new genus of erotylid beetles (Erotylidae: Pharaxonothinae) inhabiting male cones of the cycad Ceratozamia (Cycadales: Zamiaceae) Author Tang, William Author Skelley, Paul Author Pérez-Farrera, Miguel Angel text Zootaxa 2018 2018-10-31 4508 2 151 178 journal article 28054 10.11646/zootaxa.4508.2.1 da79b654-bd50-4715-8ed3-80fa7a00d469 1175-5326 2606934 5E2BC894-1919-4F63-8EF5-BAAC91913388 Key to genera of Pharaxonothinae (modified from Leschen 2003 and Xu et al. 2015 ) 1 Antennal club of 2 antennomeres.......................................................... Setariola Jacobson - Antennal club of 3 antennomeres......................................................................... 2 2 Pronotal callosities present; vestiture of scale-like setae................................ Leucohimatium Rosenhauer - Pronotal callosities absent; vestiture of simple setae.......................................................... 3 3 Eyes small, ventrally not encroaching upon gular area, distance between eye and lateral corner of mentum about equal to length of eye in ventral view; not associated with cycads...................................................... 4 - Eyes large, ventrally encroaching upon gular area, distance between eye and lateral corner of mentum <1/2 eye length in ventral view; most associated with cycads..................................................................... 5 4 Dorsal punctation strong and impressed.................................................... Henoticonus Reitter - Dorsal punctation weak and not strongly impressed......................................... Loberogosmus Reitter 5 Submental-gular suture visible as depressed darkened area, often with row of punctures and setae; anterior pronotal margin of most with fine, distinct, complete marginal line; male genitalia with median lobe and tegmen laterally flattened and slightly to strongly twisted (at least basally); spiculum gastrale asymmetrical; Asia............... Cycadophila Xu, Tang & Skelley - Submental-gular suture apparently lacking, surface flattened; anterior pronotal margin lacking marginal line, some with row of fine punctures; male genitalia with median lobe cylindrical and tegmen dorsoventrally or laterally flattened, but usually not twisted; spiculum gastrale symmetrical; Americas............................................................ 6 6 Lateral pronotal carina thicker anteriorly in lateral view ( Figs. 1 D–E); head with supraocular lines and elytra lacking basal bead; stridulatory file at base of head separated by distance ~ 0.5× width of the scutellar shield ( Fig. 1B ); tarsomeres I–III distinctly widened ( Figs. 4 D–E, 8F), metatarsomere I width ± length; male genitalia with median lobe arched in lateral view; flagellum length Ĺ length of median lobe and not coiled ( Figs. 4G , 5F , 6H , 7G , 8G ); Mexico and probably Belize , Guatemala and Honduras on Ceratozamia ............................................................ Ceratophila , new genus - Lateral pronotal carina narrow, same thickness along entire length in lateral view ( Fig. 1C ); head without supraocular lines and elytra with distinct basal bead ( Fig. 1A; P . esperanzae Chaves and Genaro is the exception); stridulatory files at base of head separated by distance ~ width of scutellar shield ( Fig. 1A ); tarsomeres I–III narrow, metatarsomere I length always distinctly greater than width; male genitalia with median lobe straight; flagellum length >> length of median lobe and coiled; widespread New World on Ceratozamia , Dioon , Microcycas , Zamia ..................................... Pharaxonotha Reitter