<document ID-DOI="10.11646/zootaxa.5263.2.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="7800978" ID-ZooBank="B07051F2-A15B-49EC-9A2F-91B9420FEA4A" approvalRequired="33" approvalRequired_for_taxonomicNames="7" approvalRequired_for_textStreams="15" approvalRequired_for_treatments="11" checkinTime="1680677567416" checkinUser="plazi" docAuthor="Gupta, Devanshu, Keith, Denis, Bhunia, Debika, Das, Priyanka, Ghosh, Joyjit &amp; Chandra, Kailash" docDate="2023" docId="BC1F87FF137CFFFAFF74FF4EFDB7BAE3" docLanguage="en" docName="zootaxa.5263.2.2.pdf" docOrigin="Zootaxa 5263 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5263.2.2" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Melolontha guttigera Sharp 1876" docType="treatment" docVersion="2" lastPageNumber="193" masterDocId="4026FF87137EFFF8FFE3FFD9FFF2B92E" masterDocTitle="Review of Melolontha Fabricius, 1775 (Coleoptera: Scarabaeidae: Melolonthinae) from India with the description of two new species" masterLastPageNumber="216" masterPageNumber="191" pageNumber="193" updateTime="1680678180874" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title>Review of Melolontha Fabricius, 1775 (Coleoptera: Scarabaeidae: Melolonthinae) from India with the description of two new species</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Gupta, Devanshu</mods:namePart>
<mods:affiliation>Zoological Survey of India, M-Block, New Alipore, Kolkata- 700053, West Bengal, India.</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Keith, Denis</mods:namePart>
<mods:affiliation>120 rue Gabriel Péri, F- 28000 Chartres, France.</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Bhunia, Debika</mods:namePart>
<mods:affiliation>Zoological Survey of India, M-Block, New Alipore, Kolkata- 700053, West Bengal, India.</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Das, Priyanka</mods:namePart>
<mods:affiliation>Zoological Survey of India, M-Block, New Alipore, Kolkata- 700053, West Bengal, India.</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ghosh, Joyjit</mods:namePart>
<mods:affiliation>Zoological Survey of India, M-Block, New Alipore, Kolkata- 700053, West Bengal, India.</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Chandra, Kailash</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0001-9076-5442</mods:nameIdentifier>
<mods:affiliation>Former Director, Zoological Survey of India, M-Block, New Alipore, Kolkata- 700053, West Bengal, India. kailash 611 @ rediffmail. com; https: // orcid. org / 0000 - 0001 - 9076 - 5442</mods:affiliation>
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<mods:title>Zootaxa</mods:title>
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<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-04-05</mods:number>
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<mods:detail type="volume">
<mods:number>5263</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>2</mods:number>
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<mods:extent unit="page">
<mods:start>191</mods:start>
<mods:end>216</mods:end>
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<mods:url>http://dx.doi.org/10.11646/zootaxa.5263.2.2</mods:url>
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<mods:identifier type="DOI">10.11646/zootaxa.5263.2.2</mods:identifier>
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<paragraph blockId="2.[151,566,151,214]" box="[151,566,151,178]" pageId="2" pageNumber="193">
<heading bold="true" box="[151,566,151,178]" fontSize="11" level="1" pageId="2" pageNumber="193" reason="1">
<taxonomicName authority="Sharp, 1876" authorityName="Sharp" authorityYear="1876" box="[151,566,151,178]" class="Insecta" family="Melolonthidae" genus="Melolontha" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="193" phylum="Arthropoda" rank="species" species="guttigera">
<emphasis bold="true" box="[151,566,151,178]" pageId="2" pageNumber="193">
<emphasis bold="true" box="[151,408,151,178]" italics="true" pageId="2" pageNumber="193">Melolontha guttigera</emphasis>
<bibRefCitation author="Sharp, D." box="[415,566,151,177]" pageId="2" pageNumber="193" pagination="65 - 90" refId="ref15828" refString="Sharp, D. (1876) Descriptions of some new species of Scarabaeidae from tropical Asia and Malaysia. Coleopterologische Hefte, 15, 65 - 90." type="journal article" year="1876">Sharp, 1876</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[151,372,187,214]" pageId="2" pageNumber="193" type="description">
<paragraph blockId="2.[151,566,151,214]" box="[151,372,187,214]" pageId="2" pageNumber="193">
(
<figureCitation box="[159,237,187,213]" captionStart="FIGURES 1–4" captionStartId="3.[151,264,1930,1954]" captionTargetBox="[167,1416,214,1888]" captionTargetId="figure-17@3.[151,1436,200,1907]" captionTargetPageId="3" captionText="FIGURES 1–4. Habitus of Melolontha species. 1, M. guttigera; 2, M. virescens; 3, M. arunachalensis new species (holotype); 4, M. lachungensis new species (holotype)." figureDoi="http://doi.org/10.5281/zenodo.7800980" httpUri="https://zenodo.org/record/7800980/files/figure.png" pageId="2" pageNumber="193">Figs. 1</figureCitation>
, 
<figureCitation box="[251,281,187,213]" captionStart="FIGURES 13–18" captionStartId="6.[151,264,1761,1785]" captionTargetBox="[152,1435,184,1728]" captionTargetId="figure-25@6.[151,1436,181,1737]" captionTargetPageId="6" captionText="FIGURES 13–18. Pygidium of Melolontha species. 13, M. guttigera; 14, M. virescens; 15, M. arunachalensis new species (holotype); 16, M. lachungensis new species (holotype); 17, M. aeneicollis; 18, M. cuprescens." figureDoi="http://doi.org/10.5281/zenodo.7800986" httpUri="https://zenodo.org/record/7800986/files/figure.png" pageId="2" pageNumber="193">13</figureCitation>
, 
<figureCitation box="[293,323,187,213]" captionStart="FIGURES 41–46" captionStartId="13.[151,264,1943,1967]" captionTargetBox="[254,1329,181,1912]" captionTargetId="figure-17@13.[252,1335,181,1920]" captionTargetPageId="13" captionText="FIGURES 41–46. Parameres of Melolontha species in dorsal view. 41, M. guttigera; 42, M. virescens; 43, M. arunachalensis new species (holotype); 44, M. lachungensis new species (holotype); 45, M. aeneicollis; 46, M. cuprescens." figureDoi="http://doi.org/10.5281/zenodo.7801000" httpUri="https://zenodo.org/record/7801000/files/figure.png" pageId="2" pageNumber="193">41</figureCitation>
, 
<figureCitation box="[335,364,187,214]" captionStart="FIGURES 53–64" captionStartId="15.[151,264,1943,1967]" captionTargetBox="[209,1358,186,1908]" captionTargetId="figure-17@15.[209,1379,181,1920]" captionTargetPageId="15" captionText="FIGURES 53–64. Parameres of Melolontha species in lateral view. 53, M. guttigera; 54, M. virescens; 55, M. arunachalensis new species (holotype); 56, M. lachungensis new species (holotype); 57, M. aeneicollis; 58, M. cuprescens; 59, M. chinensis; 60, M. nepalensis; 61, M. carinata; 62, M. indica; 63, M. flabellata; 64, M. furcicauda." figureDoi="http://doi.org/10.5281/zenodo.7801008" httpUri="https://zenodo.org/record/7801008/files/figure.png" pageId="2" pageNumber="193">53</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="193" type="reference_group">
<paragraph blockId="2.[151,1436,255,433]" pageId="2" pageNumber="193">
<treatmentCitationGroup pageId="2" pageNumber="193">
<treatmentCitation author="Sharp, D." box="[151,538,255,280]" page="87" pageId="2" pageNumber="193" year="1876">
<taxonomicName authority="Sharp 1876: 87" authorityName="Sharp" authorityPageNumber="87" authorityYear="1876" box="[151,538,255,280]" class="Insecta" family="Melolonthidae" genus="Melolontha" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="193" phylum="Arthropoda" rank="species" species="guttigera">
<emphasis box="[151,370,256,280]" italics="true" pageId="2" pageNumber="193">Melolontha guttigera</emphasis>
<bibRefCitation author="Sharp, D." box="[378,538,255,280]" pageId="2" pageNumber="193" pagination="65 - 90" refId="ref15828" refString="Sharp, D. (1876) Descriptions of some new species of Scarabaeidae from tropical Asia and Malaysia. Coleopterologische Hefte, 15, 65 - 90." type="journal article" year="1876">Sharp 1876: 87</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(type locality: 
<collectingCountry box="[699,755,256,280]" name="India" pageId="2" pageNumber="193">India</collectingCountry>
); 
<treatmentCitation author="Barlow, E." box="[775,963,256,280]" page="270" pageId="2" pageNumber="193" year="1899">
<bibRefCitation author="Barlow, E." box="[775,963,256,280]" pageId="2" pageNumber="193" pagination="234 - 273" refId="ref13039" refString="Barlow, E. (1899) List of the Melolonthini contained in the collection of the Indian Museum. Indian Museum Notes, 4 (5), 234 - 273." type="journal article" year="1899">Barlow 1899: 270</bibRefCitation>
</treatmentCitation>
(catalogue); 
<treatmentCitation author="Arrow, G. J." box="[1100,1291,256,280]" page="192" pageId="2" pageNumber="193" year="1913">
<bibRefCitation author="Arrow, G. J." box="[1100,1291,256,280]" pageId="2" pageNumber="193" pagination="191 - 196" refId="ref12795" refString="Arrow, G. J. (1913 a) Coleoptera IV: Lamellicornia. Records of the Indian Museum, 8, 191 - 196. https: // doi. org / 10.26515 / rzsi / v 8 / i 3 / 1913 / 163144" type="journal article" year="1913">Arrow 1913a: 192</bibRefCitation>
</treatmentCitation>
(comment on placement of genus and sexual dimorphism); 
<treatmentCitation author="Arrow, G. J." box="[655,840,286,310]" page="401" pageId="2" pageNumber="193" year="1913">
<bibRefCitation author="Arrow, G. J." box="[655,840,286,310]" pageId="2" pageNumber="193" pagination="394 - 408" refId="ref12842" refString="Arrow, G. J. (1913 b) Notes on the Lamellicorn Coleoptera of Japan and descriptions of a few new species. Annals and Magazine of Natural History, Series 8, 12 (70), 394 - 408. https: // doi. org / 10.1080 / 00222931308693416" type="journal article" year="1913">Arrow 1913b: 401</bibRefCitation>
</treatmentCitation>
(comment on placement of genus and sexual dimorphism); 
<treatmentCitation author="Bezdek, A." box="[199,388,317,341]" page="195" pageId="2" pageNumber="193" year="2006">
<bibRefCitation author="Bezdek, A." box="[199,388,317,341]" pageId="2" pageNumber="193" pagination="181 - 228" refId="ref13103" refString="Bezdek, A. (2006) Scarabaeidae: Melolonthinae. In: Lobl, I. &amp; Smetana, S. (Eds.), Catalogue of Palaearctic Coleoptera 3. Apollo Books, Stenstrup and Denmark, pp. 181 - 228." type="book chapter" year="2006">Bezděk 2006: 195</bibRefCitation>
</treatmentCitation>
(catalogue); 
<treatmentCitation author="Li, C. - L. &amp; Yang, P. - S. &amp; Wang, C. - C." box="[526,725,317,341]" page="351" pageId="2" pageNumber="193" year="2010">
<bibRefCitation author="Li, C. - L. &amp; Yang, P. - S. &amp; Wang, C. - C." box="[526,725,317,341]" pageId="2" pageNumber="193" pagination="341 - 359" refId="ref14954" refString="Li, C. - L., Yang, P. - S. &amp; Wang, C. - C. (2010) Revision of the Melolontha guttigera group with a key and an annotated checklist of the East and South-East Asian Melolontha groups. Annals of the Entomological Society of America, 103 (3), 341 - 359. https: // doi. org / 10.1603 / AN 09088" type="journal article" year="2010">
Li 
<emphasis box="[557,611,317,341]" italics="true" pageId="2" pageNumber="193">et al.</emphasis>
2010: 351
</bibRefCitation>
</treatmentCitation>
, figs., 24, 24, 25, 32, 37, 42, 47, 52, 57, 61 (
<typeStatus box="[1201,1296,317,341]" pageId="2" pageNumber="193">lectotype</typeStatus>
designation); Chandra &amp; Gupta 2012: 890, fig. 2A (distribution); 
<treatmentCitation author="Bezdek, A." box="[724,908,347,372]" page="231" pageId="2" pageNumber="193" year="2016">
<bibRefCitation author="Bezdek, A." box="[724,908,347,372]" pageId="2" pageNumber="193" pagination="226 - 236" refId="ref13149" refString="Bezdek, A. (2016) Melolonthini. In: Lobl, I. &amp; Lobl, D. (Eds.), Catalogue of Palaearctic Coleoptera 3 Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea. Revised and updated edition. Brill, Leiden and Boston, pp. 226 - 236." type="book chapter" year="2016">Bezděk 2016: 231</bibRefCitation>
</treatmentCitation>
(catalogue).
</treatmentCitationGroup>
</paragraph>
<paragraph blockId="2.[151,1436,255,433]" box="[151,1044,378,402]" pageId="2" pageNumber="193">
<treatmentCitationGroup box="[151,1044,378,402]" pageId="2" pageNumber="193">
<treatmentCitation author="Nonfried, A. F." box="[151,600,378,402]" page="285" pageId="2" pageNumber="193" year="1892">
<taxonomicName authority=": Nonfried 1892: 285" authorityName=": Nonfried" authorityPageNumber="285" authorityYear="1892" box="[151,600,378,402]" class="Insecta" family="Melolonthidae" genus="Hoplosternus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="193" phylum="Arthropoda" rank="species" species="guttigera">
<emphasis box="[151,387,378,402]" italics="true" pageId="2" pageNumber="193">Hoplosternus guttigera</emphasis>
: 
<bibRefCitation author="Nonfried, A. F." box="[400,600,378,402]" pageId="2" pageNumber="193" pagination="249 - 290" refId="ref15168" refString="Nonfried, A. F. (1892) Verzeichniss der seit 1871 neu beschriebenen Glaphyriden, Melolonthiden und Euchiriden. Berliner entomologische Zeitschrift, 37 (3), 249 - 290. https: // doi. org / 10.1002 / mmnd. 18920370302" type="journal article" year="1892">Nonfried 1892: 285</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(checklist); 
<treatmentCitation author="Brenske, E." box="[727,918,378,402]" page="154" pageId="2" pageNumber="193" year="1900">
<bibRefCitation author="Brenske, E." box="[727,918,378,402]" pageId="2" pageNumber="193" pagination="141 - 155" refId="ref13431" refString="Brenske, E. (1900) Contribution a letude de la faune de Sumatra (Cote ouest-Vice-residence de Painan) VIII Melolonthiden gesammelt von J-L Weyers. Memoires de la Societe Entomologique de Belgique, 7, 141 - 155." type="journal article" year="1900">Brenske 1900: 154</bibRefCitation>
</treatmentCitation>
(diagnosis).
</treatmentCitationGroup>
</paragraph>
<paragraph blockId="2.[151,1436,255,433]" box="[151,1348,409,433]" pageId="2" pageNumber="193">
<treatmentCitationGroup box="[151,1348,409,433]" pageId="2" pageNumber="193">
<treatmentCitation author="Dalla Torre, K. W. von" box="[151,611,409,433]" page="264" pageId="2" pageNumber="193" year="1912">
<taxonomicName authority=": Dalla Torre 1912: 264" authorityName="Dalla Torre" authorityPageNumber="264" authorityYear="1912" box="[151,611,409,433]" class="Insecta" family="Melolonthidae" genus="Hoplosternus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="193" phylum="Arthropoda" rank="species" species="guttiger">
<emphasis box="[151,374,409,433]" italics="true" pageId="2" pageNumber="193">Hoplosternus guttiger</emphasis>
: 
<bibRefCitation author="Dalla Torre, K. W. von" box="[388,611,409,433]" pageId="2" pageNumber="193" pagination="1 - 290" refId="ref14005" refString="Dalla Torre, K. W. von (1912) Scarabaeidae: Melolonthinae I. In: Junk, W. &amp; Schenkling, S. (Eds.), Coleopterorum Catalogus. Pars 49. W. Junk, Berlin, pp. 1 - 290." type="book chapter" year="1912">Dalla Torre 1912: 264</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(catalogue, misspelling); 
<treatmentCitation author="Sabatinelli, G." box="[874,1091,409,433]" page="613" pageId="2" pageNumber="193" year="1992">
<bibRefCitation author="Sabatinelli, G." box="[874,1091,409,433]" pageId="2" pageNumber="193" pagination="593 - 636" refId="ref15648" refString="Sabatinelli, G. (1992) Catalogue of the Scarabaeoidea Melolonthidae from Himalaya and North India: Macrodactylini, Hopliini, Diplotaxini, Melolonthini and Sericini (Coleoptera). Memorie della Societa Entomologica Italiana, 71 (2), 593 - 636." type="journal article" year="1992">Sabatinelli 1992: 613</bibRefCitation>
</treatmentCitation>
(catalogue, misspelling).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="193" type="materials_examined">
<paragraph blockId="2.[151,1437,479,973]" pageId="2" pageNumber="193">
<emphasis bold="true" box="[151,376,479,505]" pageId="2" pageNumber="193">Material examined</emphasis>
. 
<materialsCitation collectingDate="1908-09-09" collectingDateMax="2018-11-03" collectingDateMin="1908-09-09" collectionCode="NZSI" collectorName="Major Bennett &amp; N. B. Jahans &amp; D. Suresh Chand &amp; Chapramari Wildlife Sanctuary &amp; Karli Line &amp; Antipoaching Tower &amp; Ashaley Camp &amp; Samsing Beat &amp; P. C. Pathania" country="India" county="Darjeeling" elevation="229" location="Neora Valley National Park" municipality="Ghum" pageId="2" pageNumber="193" specimenCount="8" specimenCount-female="8" stateProvince="West Bengal">
<emphasis bold="true" box="[389,612,479,505]" pageId="2" pageNumber="193">
<collectingCountry box="[389,454,479,505]" name="India" pageId="2" pageNumber="193">India</collectingCountry>
, 
<collectingRegion box="[465,612,479,505]" country="India" name="West Bengal" pageId="2" pageNumber="193">West Bengal</collectingRegion>
</emphasis>
: 
<collectingCounty box="[626,747,479,505]" pageId="2" pageNumber="193">Darjeeling</collectingCounty>
, 
<collectingMunicipality box="[757,830,479,505]" pageId="2" pageNumber="193">Ghum</collectingMunicipality>
, 
<quantity box="[840,907,479,506]" metricMagnitude="2" metricUnit="m" metricValue="2.286" pageId="2" pageNumber="193" unit="ft" value="750.0">
<elevation box="[840,907,479,506]" metricMagnitude="2" metricUnit="m" metricValue="2.286" pageId="2" pageNumber="193" unit="ft" value="750.0">750ft.</elevation>
</quantity>
<date box="[912,1034,479,505]" pageId="2" pageNumber="193" value="1908-09-09">
<collectingDate box="[912,1034,479,505]" pageId="2" pageNumber="193" value="1908-09-09">09.ix.1908</collectingDate>
</date>
, 
<specimenCount box="[1045,1080,479,505]" pageId="2" pageNumber="193" type="female">1♀</specimenCount>
, Mus. Coll. [
<collectionCode box="[1223,1293,479,505]" country="India" httpUri="http://grbio.org/cool/7mhb-r8q3" name="Zoological Survey of India, National Zoological Collection" pageId="2" pageNumber="193">NZSI</collectionCode>
]; Darjeeling, Ghum, 
<quantity box="[237,303,515,542]" metricMagnitude="2" metricUnit="m" metricValue="2.286" pageId="2" pageNumber="193" unit="ft" value="750.0">
<elevation box="[237,303,515,542]" metricMagnitude="2" metricUnit="m" metricValue="2.286" pageId="2" pageNumber="193" unit="ft" value="750.0">750ft.</elevation>
</quantity>
<date box="[311,434,515,541]" pageId="2" pageNumber="193" value="1908-09-18">
<collectingDate box="[311,434,515,541]" pageId="2" pageNumber="193" value="1908-09-18">18.ix.1908</collectingDate>
</date>
, 1♁, 
<specimenCount box="[497,533,515,541]" pageId="2" pageNumber="193" type="female">2♀</specimenCount>
, leg. 
<collectorName box="[597,761,515,542]" pageId="2" pageNumber="193">Major Bennett</collectorName>
[
<collectionCode box="[777,847,515,541]" country="India" httpUri="http://grbio.org/cool/7mhb-r8q3" name="Zoological Survey of India, National Zoological Collection" pageId="2" pageNumber="193">NZSI</collectionCode>
]; Kurseong, 
<quantity box="[991,1089,515,542]" metricMagnitude="0" metricUnit="m" metricValue="1.524" pageId="2" pageNumber="193" unit="ft" value="5.0">
<elevation box="[991,1089,515,542]" metricMagnitude="0" metricUnit="m" metricValue="1.524" pageId="2" pageNumber="193" unit="ft" value="5.0">5,000 ft.</elevation>
</quantity>
, 1♁, leg. 
<collectorName box="[1203,1339,515,542]" pageId="2" pageNumber="193">N.B. Jahans</collectorName>
[
<collectionCode box="[1354,1424,515,541]" country="India" httpUri="http://grbio.org/cool/7mhb-r8q3" name="Zoological Survey of India, National Zoological Collection" pageId="2" pageNumber="193">NZSI</collectionCode>
]; Darjeeling, Mahananda Wildlife Sanctuary, Latpanchar Beat Office, 26.53.463N, 88.23.958E, 
<quantity box="[1221,1297,551,577]" metricMagnitude="3" metricUnit="m" metricValue="1.1161775999999999" pageId="2" pageNumber="193" unit="ft" value="3662.0">
<elevation box="[1221,1297,551,577]" metricMagnitude="3" metricUnit="m" metricValue="1.1161775999999999" pageId="2" pageNumber="193" unit="ft" value="3662.0">3662ft</elevation>
</quantity>
, 
<date box="[1309,1432,551,577]" pageId="2" pageNumber="193" value="2017-11-08">
<collectingDate box="[1309,1432,551,577]" pageId="2" pageNumber="193" value="2017-11-08">08.xi.2017</collectingDate>
</date>
, 3♁, 
<specimenCount box="[207,243,587,613]" pageId="2" pageNumber="193" type="female">2♀</specimenCount>
, leg. 
<collectorName box="[317,520,587,614]" pageId="2" pageNumber="193">D. Suresh Chand</collectorName>
[
<collectionCode box="[540,610,587,613]" country="India" httpUri="http://grbio.org/cool/7mhb-r8q3" name="Zoological Survey of India, National Zoological Collection" pageId="2" pageNumber="193">NZSI</collectionCode>
]; Darjeeling, Mahananda Wildlife Sanctuary, Latpanchar Beat Office, 26.53.463N, 88.23.958E, 
<quantity box="[438,526,623,649]" metricMagnitude="3" metricUnit="m" metricValue="1.1161775999999999" pageId="2" pageNumber="193" unit="ft" value="3662.0">
<elevation box="[438,526,623,649]" metricMagnitude="3" metricUnit="m" metricValue="1.1161775999999999" pageId="2" pageNumber="193" unit="ft" value="3662.0">3662 ft.</elevation>
</quantity>
, 
<date box="[536,644,623,649]" pageId="2" pageNumber="193" value="2018-11-03">
<collectingDate box="[536,644,623,649]" pageId="2" pageNumber="193" value="2018-11-03">3.xi.2018</collectingDate>
</date>
, 2♁, 
<specimenCount box="[702,738,623,649]" pageId="2" pageNumber="193" type="female">3♀</specimenCount>
, leg. 
<collectorName box="[796,984,623,650]" pageId="2" pageNumber="193">D. Suresh Chand</collectorName>
[
<collectionCode box="[996,1066,623,649]" country="India" httpUri="http://grbio.org/cool/7mhb-r8q3" name="Zoological Survey of India, National Zoological Collection" pageId="2" pageNumber="193">NZSI</collectionCode>
]; 
<collectorName box="[1084,1432,623,649]" pageId="2" pageNumber="193">Chapramari Wildlife Sanctuary</collectorName>
, 
<collectorName box="[151,269,659,685]" pageId="2" pageNumber="193">Karli Line</collectorName>
, 
<collectorName box="[278,507,659,686]" pageId="2" pageNumber="193">Antipoaching Tower</collectorName>
<quantity box="[513,607,659,685]" metricMagnitude="-1" metricUnit="m" metricValue="5.306568" pageId="2" pageNumber="193" unit="ft" value="1.741">
<elevation box="[513,607,659,685]" metricMagnitude="-1" metricUnit="m" metricValue="5.306568" pageId="2" pageNumber="193" unit="ft" value="1.741">1, 741ft.</elevation>
</quantity>
<date box="[613,727,659,685]" pageId="2" pageNumber="193" value="2018-10-31">
<collectingDate box="[613,727,659,685]" pageId="2" pageNumber="193" value="2018-10-31">31.x.2018</collectingDate>
</date>
, 1♁, leg. 
<collectorName box="[833,1022,659,686]" pageId="2" pageNumber="193">D. Suresh Chand</collectorName>
[
<collectionCode box="[1035,1105,659,685]" country="India" httpUri="http://grbio.org/cool/7mhb-r8q3" name="Zoological Survey of India, National Zoological Collection" pageId="2" pageNumber="193">NZSI</collectionCode>
]; 
<location LSID="urn:lsid:plazi:treatment:BC1F87FF137CFFFAFF74FF4EFDB7BAE3:31696032137CFFFAFB80FD4DFA6ABB83" box="[1123,1432,659,686]" country="India" county="Darjeeling" municipality="Ghum" name="Neora Valley National Park" pageId="2" pageNumber="193" stateProvince="West Bengal">Neora Valley National Park</location>
, 
<collectorName box="[151,322,695,721]" pageId="2" pageNumber="193">Ashaley Camp</collectorName>
, 
<collectorName box="[336,496,695,722]" pageId="2" pageNumber="193">Samsing Beat</collectorName>
, 
<date box="[510,632,695,721]" pageId="2" pageNumber="193" value="2018-09-22">
<collectingDate box="[510,632,695,721]" pageId="2" pageNumber="193" value="2018-09-22">22.ix.2018</collectingDate>
</date>
, 1♁, leg. 
<collectorName box="[749,899,695,721]" pageId="2" pageNumber="193">P.C. Pathania</collectorName>
[
<collectionCode box="[916,987,695,721]" country="India" httpUri="http://grbio.org/cool/7mhb-r8q3" name="Zoological Survey of India, National Zoological Collection" pageId="2" pageNumber="193">NZSI</collectionCode>
]
</materialsCitation>
. 
<materialsCitation collectingDate="1998-09-07" collectingDateMax="2015-10-21" collectingDateMin="1998-09-07" collectionCode="NZSI" collectorName="P. T. Bhutia &amp; A. R. Lahiri &amp; S. K. Mandal &amp; Dhriti Banerjee &amp; M. Sil &amp; S. D. Gurumayum" country="India" county="Itanagar" elevation="216" location="Kameng" municipality="Complex" pageId="2" pageNumber="193" specimenCount="2" specimenCount-female="1" stateProvince="Arunachal Pradesh">
<emphasis bold="true" box="[1007,1321,695,721]" pageId="2" pageNumber="193">
<collectingCountry box="[1007,1072,695,721]" name="India" pageId="2" pageNumber="193">India</collectingCountry>
, 
<collectingRegion box="[1086,1321,695,721]" country="India" name="Arunachal Pradesh" pageId="2" pageNumber="193">Arunachal Pradesh</collectingRegion>
</emphasis>
: 
<collectingCounty box="[1339,1433,696,722]" pageId="2" pageNumber="193">Itanagar</collectingCounty>
, ZSI 
<collectingMunicipality box="[200,306,731,757]" pageId="2" pageNumber="193">Complex</collectingMunicipality>
, 
<specimenCount box="[316,372,731,758]" pageId="2" pageNumber="193" type="generic">1 ex.</specimenCount>
, leg. 
<collectorName box="[432,555,731,758]" pageId="2" pageNumber="193">P.T. Bhutia</collectorName>
[
<collectionCode box="[568,638,731,757]" country="India" httpUri="http://grbio.org/cool/7mhb-r8q3" name="Zoological Survey of India, National Zoological Collection" pageId="2" pageNumber="193">NZSI</collectionCode>
]; 
<location LSID="urn:lsid:plazi:treatment:BC1F87FF137CFFFAFF74FF4EFDB7BAE3:31696032137CFFFAFD72FD05FD01BBD8" box="[657,755,732,758]" country="India" county="Itanagar" municipality="Complex" name="Kameng" pageId="2" pageNumber="193" stateProvince="Arunachal Pradesh">Kameng</location>
, 
<location LSID="urn:lsid:plazi:treatment:BC1F87FF137CFFFAFF74FF4EFDB7BAE3:31696032137CFFFAFD1EFD02FC71BBDB" box="[765,899,731,757]" country="India" county="Itanagar" municipality="Complex" name="Bhalulpong" pageId="2" pageNumber="193" stateProvince="Arunachal Pradesh">Bhalulpong</location>
, 
<date box="[909,1018,731,757]" pageId="2" pageNumber="193" value="1998-09-07">
<collectingDate box="[909,1018,731,757]" pageId="2" pageNumber="193" value="1998-09-07">7.ix.1998</collectingDate>
</date>
, 1♁, leg. 
<collectorName box="[1124,1251,731,757]" pageId="2" pageNumber="193">A.R. Lahiri</collectorName>
[
<collectionCode box="[1265,1335,731,757]" country="India" httpUri="http://grbio.org/cool/7mhb-r8q3" name="Zoological Survey of India, National Zoological Collection" pageId="2" pageNumber="193">NZSI</collectionCode>
]; 
<location LSID="urn:lsid:plazi:treatment:BC1F87FF137CFFFAFF74FF4EFDB7BAE3:31696032137CFFFAFAA9FD02FF12BA37" country="India" county="Itanagar" municipality="Complex" name="Dibang Valley" pageId="2" pageNumber="193" stateProvince="Arunachal Pradesh">Dibang Valley</location>
, 
<location LSID="urn:lsid:plazi:treatment:BC1F87FF137CFFFAFF74FF4EFDB7BAE3:31696032137CFFFAFF08FD26FEC0BA37" box="[235,306,767,793]" country="India" county="Itanagar" municipality="Complex" name="Roing" pageId="2" pageNumber="193" stateProvince="Arunachal Pradesh">Roing</location>
, 
<location LSID="urn:lsid:plazi:treatment:BC1F87FF137CFFFAFF74FF4EFDB7BAE3:31696032137CFFFAFEDEFD26FEADBA37" box="[317,351,767,793]" country="India" county="Itanagar" municipality="Complex" name="Jia" pageId="2" pageNumber="193" stateProvince="Arunachal Pradesh">Jia</location>
, 
<date box="[362,470,767,793]" pageId="2" pageNumber="193" value="2000-01-26">
<collectingDate box="[362,470,767,793]" pageId="2" pageNumber="193" value="2000-01-26">26.i.2000</collectingDate>
</date>
, 1♁, leg. 
<collectorName box="[578,720,767,793]" pageId="2" pageNumber="193">S.K. Mandal</collectorName>
[
<collectionCode box="[733,803,767,793]" country="India" httpUri="http://grbio.org/cool/7mhb-r8q3" name="Zoological Survey of India, National Zoological Collection" pageId="2" pageNumber="193">NZSI</collectionCode>
]; 
<location LSID="urn:lsid:plazi:treatment:BC1F87FF137CFFFAFF74FF4EFDB7BAE3:31696032137CFFFAFCD5FD26FB96BA34" box="[822,1124,767,794]" country="India" county="Itanagar" municipality="Complex" name="Bhalukpong Circuit House" pageId="2" pageNumber="193" stateProvince="Arunachal Pradesh">Bhalukpong Circuit House</location>
, 
<quantity box="[1135,1202,767,793]" metricMagnitude="2" metricUnit="m" metricValue="2.16" pageId="2" pageNumber="193" unit="m" value="216.0">
<elevation box="[1135,1202,767,793]" metricMagnitude="2" metricUnit="m" metricValue="2.16" pageId="2" pageNumber="193" unit="m" value="216.0">216m</elevation>
</quantity>
, 
<date box="[1213,1335,767,793]" pageId="2" pageNumber="193" value="2001-09-30">
<collectingDate box="[1213,1335,767,793]" pageId="2" pageNumber="193" value="2001-09-30">30.ix.2001</collectingDate>
</date>
, 1♁, leg. 
<collectorName box="[151,324,803,829]" pageId="2" pageNumber="193">Dhriti Banerjee</collectorName>
[
<collectionCode box="[336,406,803,829]" country="India" httpUri="http://grbio.org/cool/7mhb-r8q3" name="Zoological Survey of India, National Zoological Collection" pageId="2" pageNumber="193">NZSI</collectionCode>
]; Domrah, 
<quantity box="[528,595,803,829]" metricMagnitude="2" metricUnit="m" metricValue="4.8" pageId="2" pageNumber="193" unit="m" value="480.0">
<elevation box="[528,595,803,829]" metricMagnitude="2" metricUnit="m" metricValue="4.8" pageId="2" pageNumber="193" unit="m" value="480.0">480m</elevation>
</quantity>
, 
<date box="[605,727,803,829]" pageId="2" pageNumber="193" value="2002-09-27">
<collectingDate box="[605,727,803,829]" pageId="2" pageNumber="193" value="2002-09-27">27.ix.2002</collectingDate>
</date>
, 
<specimenCount box="[738,773,803,829]" pageId="2" pageNumber="193" type="female">1♀</specimenCount>
, leg. 
<collectorName box="[833,901,803,830]" pageId="2" pageNumber="193">M. Sil</collectorName>
[
<collectionCode box="[914,984,803,829]" country="India" httpUri="http://grbio.org/cool/7mhb-r8q3" name="Zoological Survey of India, National Zoological Collection" pageId="2" pageNumber="193">NZSI</collectionCode>
]; 
<location LSID="urn:lsid:plazi:treatment:BC1F87FF137CFFFAFF74FF4EFDB7BAE3:31696032137CFFFAFC09FCFDFA6ABA10" box="[1002,1432,803,830]" country="India" county="Itanagar" municipality="Complex" name="Lower Dibang Valley Patrolling Route" pageId="2" pageNumber="193" stateProvince="Arunachal Pradesh">Lower Dibang Valley Patrolling Route</location>
, 
<date box="[151,265,839,866]" pageId="2" pageNumber="193" value="2015-10-21">
<collectingDate box="[151,265,839,866]" pageId="2" pageNumber="193" value="2015-10-21">21.x.2015</collectingDate>
</date>
, 1♁, leg. 
<collectorName box="[374,573,839,865]" pageId="2" pageNumber="193">S.D. Gurumayum</collectorName>
[
<collectionCode box="[586,657,839,865]" country="India" httpUri="http://grbio.org/cool/7mhb-r8q3" name="Zoological Survey of India, National Zoological Collection" pageId="2" pageNumber="193">NZSI</collectionCode>
]
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection box="[199,757,875,902]" pageId="2" pageNumber="193" type="diagnosis">
<paragraph blockId="2.[151,1437,479,973]" box="[199,757,875,902]" pageId="2" pageNumber="193">
<emphasis bold="true" box="[199,322,875,901]" pageId="2" pageNumber="193">Diagnosis.</emphasis>
Refer to the key to the species groups.
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="193" type="distribution">
<paragraph blockId="2.[151,1437,479,973]" pageId="2" pageNumber="193">
<emphasis bold="true" box="[199,353,911,937]" pageId="2" pageNumber="193">Distribution.</emphasis>
<collectingCountry box="[362,424,911,937]" name="India" pageId="2" pageNumber="193">India</collectingCountry>
: 
<collectingRegion box="[434,649,911,937]" country="India" name="Arunachal Pradesh" pageId="2" pageNumber="193">Arunachal Pradesh</collectingRegion>
(Chandra &amp; Gupta 2012), 
<collectingRegion box="[961,1042,911,937]" country="India" name="Sikkim" pageId="2" pageNumber="193">Sikkim</collectingRegion>
(
<bibRefCitation author="Li, C. - L. &amp; Yang, P. - S. &amp; Wang, C. - C." box="[1059,1222,911,937]" pageId="2" pageNumber="193" pagination="341 - 359" refId="ref14954" refString="Li, C. - L., Yang, P. - S. &amp; Wang, C. - C. (2010) Revision of the Melolontha guttigera group with a key and an annotated checklist of the East and South-East Asian Melolontha groups. Annals of the Entomological Society of America, 103 (3), 341 - 359. https: // doi. org / 10.1603 / AN 09088" type="journal article" year="2010">
Li 
<emphasis box="[1095,1153,911,937]" italics="true" pageId="2" pageNumber="193">et al.</emphasis>
2010
</bibRefCitation>
), and 
<collectingRegion box="[1292,1436,911,938]" country="India" name="West Bengal" pageId="2" pageNumber="193">West Bengal</collectingRegion>
(
<bibRefCitation author="Barlow, E." box="[159,310,947,973]" pageId="2" pageNumber="193" pagination="234 - 273" refId="ref13039" refString="Barlow, E. (1899) List of the Melolonthini contained in the collection of the Indian Museum. Indian Museum Notes, 4 (5), 234 - 273." type="journal article" year="1899">Barlow 1899</bibRefCitation>
); 
<collectingCountry box="[330,397,947,973]" name="Nepal" pageId="2" pageNumber="193">Nepal</collectingCountry>
(
<bibRefCitation author="Li, C. - L. &amp; Yang, P. - S. &amp; Wang, C. - C." box="[412,569,947,973]" pageId="2" pageNumber="193" pagination="341 - 359" refId="ref14954" refString="Li, C. - L., Yang, P. - S. &amp; Wang, C. - C. (2010) Revision of the Melolontha guttigera group with a key and an annotated checklist of the East and South-East Asian Melolontha groups. Annals of the Entomological Society of America, 103 (3), 341 - 359. https: // doi. org / 10.1603 / AN 09088" type="journal article" year="2010">
Li 
<emphasis box="[445,501,947,973]" italics="true" pageId="2" pageNumber="193">et al.</emphasis>
2010
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>