Macrodactylini (Coleoptera, Scarabaeidae, Melolonthinae): primary types of type species and taxonomic changes to the generic classification Author Fuhrmann, Juares Author Vaz-De-Mello, Fernando Z. text European Journal of Taxonomy 2017 2017-09-13 350 1 71 journal article 22059 10.5852/ejt.2017.350 52e4ae01-f51f-4c4a-8a14-9fb5b17788d6 2118-9773 3832682 8D14DBDE-AD13-445B-B2D0-221F19FC7C37 Pseudoserica Guérin-Méneville, 1838 , revalidated Pseudoserica Guérin-Méneville, 1838: 86 . Gama Blanchard, 1850: 118 , syn. nov . Type species: Gama grandicornis Blanchard, 1850 (designation: Lacordaire 1856: 265 ). Harpodactyla Burmeister, 1855: 17 , syn. nov . Type species: Harpodactyla grandicornis Burmeister, 1855 (designation: Evans 2003: 268 ). Pachylotoma Blanchard, 1850: 121 , syn. nov . Type species: Pachylotoma viridis Blanchard, 1850 (designation: monotypy). Pseudoserica – Blanchard 1850: 128 (synonymy with Plectris ). Harpodactyla – Gemminger & Harold 1869: 1411 (synonymy with Gama ). Pachylotoma – Katovich 2008: 6 (synonymy with Gama ). Type species Serica marmorea Guérin-Méneville, 1831 (designation: monotypy). Diagnosis Clypeal posterior angle partially covering the canthus ( Fig. 13A ); pronotal anterior margin beaded, posterior margin not beaded and with two acute projections over the anterior elytron–scutellum contact, posterior angle acute ( Fig. 13A ); prosternum anteriorly concave (similar to Fig. 12K ) or an anterior longitudinal carina; protibia with 1–3 external teeth and with or without spur; elytron with posterior margin not beaded; male mesotarsomere I with an internal hook-like spine ( Fig. 13B ); abdomen with intersegmental membrane VII–VIII concealed. Remarks The genus is recognized by having the posterior pronotal margin with two lateromedial teeth which project over the anterior elytron–scutellum contact. In addition, the male mesotarsomere I has a hooklike spine ( Fig. 13B ), that also occurs in Mallotarsus ( Fig. 13C ) and part of Dicrania (e.g., D . ebenina ). The present study revalidates Pseudoserica , formerly a synonym of Plectris , based on the following characters (opposition to Plectris ): posterior pronotal margin with two lateromedial teeth over the anterior elytron–scutellum contact (margin straight); male mesotarsomere I with a hook-like spine (without spines); internal area of metatarsomere V without spine-like setae (with spine-like setae).