Dendropaemon Perty, 1830: taxonomy, systematics and phylogeny of the morphologically most derived phanaeine genus (Coleoptera: Scarabaeidae, Scarabaeinae, Phanaeini) Author François Génier Author Patrick Arnaud text Zootaxa 2016 4099 1 journal volume 10.11646/zootaxa.4099.1.1 5d7eab55-66e2-43ed-b764-bac0c6e5b6d6 1175-5326 268274 B73C1BE1-346A-4C89-86B2-51CDACE92877 Dendropaemon (Titthopaemon) Génier & Arnaud , new subgenus Type species : Dendropaemon denticollis Felsche, 1909 ; monotypy. Description . Size small. Habitus elongated, parallel sided; with metallic sheen on head, pronotum and elytra. Body strongly compressed dorsoventrally. Clypeal edge distinctly emarginate on external side of each clypeal tooth; clypeal teeth acutely angular. Pronotum with fine punctures on disc; anterior margin flat lateral to eyes, flat surface produced into a sharp tubercle laterally; lateral fossae rounded and simple. Elytral base lacking margin. Meso and metatarsi similar in shape, three segmented, first segment approximately two times as long as wide at apex, last segment spiniformly produced internally, with setae apically. Etymology . “ Titthos ” (nipple, teat) a Greek name pertaining to the tubercle of the pronotal anterior margin which is presenting the aspect of a nipple in the species included in the subgenus, with the suffix “paemon” to keep the naming scheme similar to the genus. Gender masculine.