Morphology-based cladistics splinters the century-old dichotomy of the pied harvestmen (Arachnida: Gonyleptoidea: Cosmetidae) Author Medrano, Miguel Author Kury, Adriano Brilhante Author Mendes, Amanda Cruz text Zoological Journal of the Linnean Society 2022 2021-08-30 195 585 672 journal article 113070 10.1093/zoolinnean/zlab043 e070d52f-d008-4f08-988a-a8b17c27418f 0024-4082 6959516 1C89556A-12CB-43B7-9B49-E02EFF1543D3 KEY TO THE SPECIES OF PROTUS 1. Coxa IV with basal thickening ( Figs 5I , 31A ) and with a lateral yellow stripe ( Fig. 31B )........................... 2 1’. Coxa IV convex and without basal thickening ( Fig. 35A, C ) and with or without yellow spots but without lateral stripe ( Figs 35F , 39C, D ) ................................................................................................... 3 2. Area II of mesotergum with two paramedian rounded buttons ( Fig. 33A, B )............................. P. distinctus 2’. Area II of mesotergum unarmed ( Fig. 31B, C ) ............................................................................ P. bolivari 3. Coxa IV with scattered yellow tubercles in the lateral region ( Fig. 39C, D ).................... P. quadripunctatus 3’. Coxa IV without scattered yellow tubercles or with a single small spot ( Figs 35F , 40B ) ....................... 4 4. DS yellow markings restricted to solid strips in lateral margins ( Fig. 40A ) ............................... P. speciosus 4’. DS yellow markings only as paramedian dots in area I, III and IV ( Fig. 35A ), or as a reticulate pattern ( Fig. 37 ), but without lateral solid strips................................................................................................... 5 5. DS scutum with reticulate pattern and anterior margin coloured ( Fig. 37 ); coxa with a single lateral yellow spot ( Fig. 37B ); stylus of the penis with a small wattle ( Fig. 38A–C )............................................ P. marllusi 5’. DS scutum only with paramedian dots in areas I, III and IV ( Fig. 35A ); coxa without any yellow marks ( Fig. 35C, F ); stylus of the penis with a large wattle ( Figs 35G, H , 36A–C ) ............................. P. insolens