Taxonomy and phylogeny of the ‘ football stars’ (Asteroidea, Sphaerasteridae) Author Gale, Andrew Scott School of the Environment, Geography and Geological Sciences, University of Portsmouth, Burnaby Building, Burnaby Road, Portsmouth PO 1 3 QL UK; & Department of Earth Sciences, Natural History Museum, Cromwell Road, London SW 7 5 BD, UK text Journal of Systematic Palaeontology 2021 2021-10-01 19 10 691 741 http://dx.doi.org/10.1080/14772019.2021.1960911 journal article 10.1080/14772019.2021.1960911 1478-0941 F8991F09-B5FB-40EF-B4CC-474D925085B8 Genus Testudinaster Hess, 1983 Type species. Testudinaster pereginus Hess, 1983 , by original designation. Diagnosis. Sphaerasterida in which the abactinal surface is made up of numerous, rather small polygonal ossicles. Actinal ossicles small, very numerous, rounded or rhombic in outline, and very thick.