The braconid parasitoids (Hymenoptera: Braconidae) associated with seeds of Dipterocarpaceae in Malaysia
Author
Achterberg, C. van
Author
Hosaka, T.
Author
Ng, Y. F.
Author
Ghani, Idris B. A.
text
Journal of Natural History
2009
2009-03-31
43
11 - 12
635
686
http://dx.doi.org/10.1080/00222930802610501
journal article
10.1080/00222930802610501
1464-5262
5216015
Bracon singaporensis
Szépligeti, 1905
(
Figure 4E–H
)
Bracon singaporensis
Szépligeti 1905
, p. 37
;
Shenefelt 1978
, p. 1539.
Variation
Length of forewing of female
2.9–3.1 mm
(of male
2.4 mm
), and of body
3.3–3.5 mm
(of male
2.7 mm
); antenna of female with 30 (1) or 31 (1) segments; ovipositor sheath about as long as metasoma and 0.50–0.53 times as long as forewing; male has posterior half of third tergite and most of fourth and fifth tergites dark brown.
Biology
The small subgenus
Ophthalmo-bracon
Tobias contains parasitoids of
Gelechiidae
(
Lepidoptera
: Gelechioidea) and it is therefore likely that
B. singaporensis
parasitizes larvae of
Gelechiidae
feeding on
Shorea
seeds. However, we have not identified any
Gelechiidae
spp., but have some
Oecophoridae
species (
Lepidoptera
: Gelechioidea;
Table 1
) as possible hosts.
Notes
Dr Jenö Papp (Budapest) examined the
holotype
and kindly informed the first author that the malar space was much shorter than the basal width of the mandible. Therefore, it is obvious that
B. singaporensis
belongs to the subgenus
Ophthalmobracon
. It has the dorsolateral carinae of first tergite present (absent in Palaearctic spp.), the middle lobe of mesoscutum densely setose, the body yellowish-brown, but the third and fourth metasomal tergites of female with a pair of dark brown spots (and also sixth tergite in male), the propodeum (except anteriorly) largely and the vertex finely granulate, the ovipositor sheath is about 0.5 times the forewing and about as long as the metasoma and the marginal cell of the forewing reaches about the apex of the forewing (
Figure 4E
). Two species included in this subgenus can be separated because they have the ovipositor sheath distinctly shorter than the metasoma and vein 1-R1 of forewing about as long as pterostigma: the Central Asian
B. (O.) nocturnus
(Tobias, 1962)
and
B. (O.) lissothorax
(Tobias, 1964)
.
Bracon singaporensis
is similar to the Central and East Palaearctic
B. (O.) ophthalmicus
Telenga, 1933
, and the Afrotropical
B. (O.) kirkpatricki
(Wilkinson, 1927)
having the ovipositor sheath about as long as the metasoma, but
B. ophthalmicus
has vein SR1 of the forewing 2.3–3.0 times as long as vein 3-SR (1.3– 1.4 times in
B. singaporensis
, 1.6–1.9 times in
B. kirkpatricki
), no dorsolateral carinae on the first tergite, the third and fourth metasomal tergites completely yellowishbrown and antenna of female with 24–28 segments (30 or 31 segments in
B. singaporensis
). In addition,
B. kirkpatricki
has the forewing membrane (except basally) and the pterostigma medially darkened.
Bracon greeni
Ashmead, 1896
, from
Sri Lanka
is similarly coloured but has the ovipositor sheath about 0.7 times as long as the metasoma, the pterostigma brown, the antenna brownish-black, third tergite about as long as the fourth tergite, the second tergite shagreened to closely punctured and with a weak median carina, vein 3-SR of the forewing about three times as long as vein r and the
type
series was reared from a host [
Laccifer albizziae
(Green)
(
Kerriidae
; Coccoidea)] on shrubs and they were not associated with the seeds of trees.
Figure 5. (A, B)
Bracon rugulifer
, female, holotype; (C–G)
Bracon nitidisoma
, female, holotype; (H, I)
Syntomernus shoreatus
, female, holotype. (A, C) Part of forewing; (B, G, I) first to third metasomal tergites, dorsal aspect; (D) head, dorsal aspect; (E, H) head, anterior aspect; (F) head, lateral aspect. Scale bars: 1.0×(A); 1.3×(B, C); 1.4×(D–G); 1.8×(H, I).
In addition, we examined
two males
from
Shorea acuminata
seeds belonging to another species with the mesoscutum largely smooth.
Material examined
One female, ‘‘W.
Malaysia
, N. Sembilan Pasoh Forest Reserve,
25 January 2002
. From
Shorea acuminata
seeds, Y.F. Ng and Hosaka’’, ‘‘C 13’’, ‘‘Collected from
S. acuminata
seed (324995)’’ (UKM)
;
two females
and
one male
, id., but C 1, C 3 or C 14 (
RMNH
)
;
two females
, id., but
12 January 2002
. no. 324995 and B 15 or B 11 (
UKM
)
;
one male
, id., but
25 January 2002
, no. 324995 and C 15 (
RMNH
)
;
one male
, id., but
10 December 2001
, from
Shorea pauciflora
seeds, no. 255519 and C 9 (
UKM
)
;
one female
, id., but
1 December 2001
, no. 255519 and C 8 (
UKM
)
;
one female
, id.,
12 January 2002
, from
Shorea macroptera
seeds, no. 215945 and C 5 (
RMNH
)
.