The genus Phyllophaga Harris (Coleoptera: Scarabaeidae: Melolonthinae) in the Colombian Andean Mountains Author Vallejo, Luís Fernando Author Wolff, Martha text Zootaxa 2013 3722 2 101 142 journal article 10.11646/zootaxa.3722.2.1 f2c46514-f165-43d3-a574-47c55b89dcd6 1175-5326 218998 29D77A16-096D-4FC1-A5B4-9EEDF2E761A7 31. Phyllophaga ( Phyllophaga ) columbiana (Blanchard, 1851) Comments. The original description was based on a female specimen from "Nouv-Grenade". Blackwelder (1944) cited it for Colombia , but there are no more references as to its distribution or taxonomic position. In order to identify this species accurately, it is necessary to study the holotype , possibly deposited in the Natural History Museum in Paris.