<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1184.106698" ID-Pensoft-Pub="1313-2970-1184-301" ID-Pensoft-UUID="283A8869D1C3500A8A3C075E14584C90" ID-ZooBank="4EB4BEE3865E4CF29F7B6477D2DEF4AF" ModsDocID="1313-2970-1184-301" checkinTime="1700745464390" checkinUser="pensoft" docAuthor="Romero-Ortiz, Catalina, Sarmiento, Carlos E. &amp; Harvey, Mark S." docDate="2023" docId="19EF511689415CF698D61E7094E9FF2D" docLanguage="en" docName="ZooKeys 1184: 301-326" docOrigin="ZooKeys 1184" docPubDate="2023-11-22" docSource="http://dx.doi.org/10.3897/zookeys.1184.106698" docTitle="Cystowithius Harvey 2004" docType="treatment" docVersion="1" id="283A8869D1C3500A8A3C075E14584C90" lastPageNumber="301" masterDocId="283A8869D1C3500A8A3C075E14584C90" masterDocTitle="A new genus and five new species of pseudoscorpions (Arachnida, Pseudoscorpiones, Withiidae) from Colombia" masterLastPageNumber="326" masterPageNumber="301" pageNumber="301" updateTime="1700745464390" updateUser="pensoft">
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<mods:title id="D99242E2584D1300187C94A94807769A">A new genus and five new species of pseudoscorpions (Arachnida, Pseudoscorpiones, Withiidae) from Colombia</mods:title>
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<mods:namePart id="A1F94F70C313B93D5E67A66C0002DE8E">Romero-Ortiz, Catalina</mods:namePart>
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<mods:affiliation id="0E988439AF1F28B5B766B25B785400A1">Laboratorio de Sistematica y Biologia Comparada de Insectos, Instituto de Ciencias Naturales, Universidad Nacional de Colombia, Bogota, Colombia</mods:affiliation>
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<mods:namePart id="4076A0EE1DECF734C9B637866743CAD8">Sarmiento, Carlos E.</mods:namePart>
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<mods:affiliation id="F25B38AA15583FCAD18A85798A7B73CD">Laboratorio de Sistematica y Biologia Comparada de Insectos, Instituto de Ciencias Naturales, Universidad Nacional de Colombia, Bogota, Colombia</mods:affiliation>
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<mods:namePart id="71A7C8EBFF82CB022023D762C2D25400">Harvey, Mark S.</mods:namePart>
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<mods:affiliation id="74D19E7C0417570CB0F70BEA90EF7A57">Collections and Research, Western Australian Museum, Welshpool, Western Australia 6106, Australia &amp; School of Biological Sciences, The University of Western Australia, Crawley, Western Australia 6009, Australia</mods:affiliation>
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Genus 
<taxonomicName id="9F29E5C26C709DD61E352F580694B8E4" ID-CoL="3Z58" LSID="19EF5116-8941-5CF6-98D6-1E7094E9FF2D" authority="Harvey, 2004" authorityName="Harvey" authorityYear="2004" class="Arachnida" family="Withiidae" genus="Cystowithius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cystowithius" order="Pseudoscorpiones" pageId="0" pageNumber="301" phylum="Arthropoda" rank="genus">Cystowithius Harvey, 2004</taxonomicName>
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<paragraph id="F40301FAF2059BFCB1E8EDAAC6A12050" pageId="0" pageNumber="301">
<taxonomicName id="85191DB5D7F96CF2B9F4CEAF30EE8938" authorityName="Harvey" authorityYear="2004" class="Arachnida" family="Withiidae" genus="Cystowithius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cystowithius" order="Pseudoscorpiones" pageId="0" pageNumber="301" phylum="Arthropoda" rank="genus">Cystowithius</taxonomicName>
<bibRefCitation id="519E5862B574C815339B3F97B3BAA055" DOI="https://doi.org/10.1636/S03-48" author="Harvey, MS" journalOrPublisher="The Journal of Arachnology" pageId="0" pageNumber="301" pagination="436 - 456" refId="B15" refString="Harvey, MS, 2004. Remarks on the New World pseudoscorpion genera Parawithius and Victorwithius, with a new genus bearing a remarkable sternal modification (Pseudo-scorpiones, Withiidae). The Journal of Arachnology 32 (3): 436 - 456, DOI: https://doi.org/10.1636/S03-48" title="Remarks on the New World pseudoscorpion genera Parawithius and Victorwithius, with a new genus bearing a remarkable sternal modification (Pseudo-scorpiones, Withiidae)." url="https://doi.org/10.1636/S03-48" volume="32" year="2004">Harvey 2004</bibRefCitation>
: 440; 
<bibRefCitation id="52A1136E77B99B080AE23E13299217EE" DOI="https://doi.org/10.1590/s1984-4689.v38.e21003" author="Garcia, F" journalOrPublisher="Zoologia" pageId="0" pageNumber="301" refId="B13" refString="Garcia, F, Romero-Ortiz, C, 2021. Cystowithius ankeri sp. nov., a new pseudoscorpion from the Central Andes of Colombia (Arachnida: Pseudoscorpiones: Withiidae). Zoologia (Curitiba) 38: e21003. https://doi.org/10.1590/s1984-4689.v38.e21003" title="Cystowithius ankeri sp. nov., a new pseudoscorpion from the Central Andes of Colombia (Arachnida: Pseudoscorpiones: Withiidae). Zoologia (Curitiba) 38: e 21003." url="https://doi.org/10.1590/s1984-4689.v38.e21003" year="2021">
<normalizedToken id="76E42D47346175A6323DEA4D15649FB6" originalValue="García">Garcia</normalizedToken>
and Romero-Ortiz 2021
</bibRefCitation>
: 2.
</paragraph>
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<paragraph id="967056A05DDC99351F8315A5C8469CD7" pageId="0" pageNumber="301">Type species.</paragraph>
<paragraph id="3BE37736BE0D9C74D1C8895757751AEA" pageId="0" pageNumber="301">
<taxonomicName id="E97E989D406F1A2A90599FB02BB588B6" authorityName="Harvey" authorityYear="2004" class="Arachnida" family="Withiidae" genus="Cystowithius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cystowithius smithersi" order="Pseudoscorpiones" pageId="0" pageNumber="301" phylum="Arthropoda" rank="species" species="smithersi">
<emphasis id="F79682C284715FB35DF140F76C7FA23B" italics="true" pageId="0" pageNumber="301">Cystowithius smithersi</emphasis>
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Harvey, 2004, by original designation.
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<subSubSection id="B805A6DA8240F509D401456E5103F390" pageId="0" pageNumber="301" type="diagnosis">
<paragraph id="B779A3FDC24BCF2F45CD0C4A7208C6AA" pageId="0" pageNumber="301">Diagnosis.</paragraph>
<paragraph id="04D497DB97FA08303BC4EB05FAA626E9" pageId="0" pageNumber="301">
See 
<bibRefCitation id="C024890ACC4506113F78D24D0A3A8F03" DOI="https://doi.org/10.1636/S03-48" author="Harvey, MS" journalOrPublisher="The Journal of Arachnology" pageId="0" pageNumber="301" pagination="436 - 456" refId="B15" refString="Harvey, MS, 2004. Remarks on the New World pseudoscorpion genera Parawithius and Victorwithius, with a new genus bearing a remarkable sternal modification (Pseudo-scorpiones, Withiidae). The Journal of Arachnology 32 (3): 436 - 456, DOI: https://doi.org/10.1636/S03-48" title="Remarks on the New World pseudoscorpion genera Parawithius and Victorwithius, with a new genus bearing a remarkable sternal modification (Pseudo-scorpiones, Withiidae)." url="https://doi.org/10.1636/S03-48" volume="32" year="2004">Harvey (2004)</bibRefCitation>
.
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<subSubSection id="317CF839CE34B078660D5B722D5EBC2E" pageId="0" pageNumber="301" type="remarks">
<paragraph id="8D042CCE29FA6CBFC40BAA7646A392A1" pageId="0" pageNumber="301">Remarks.</paragraph>
<paragraph id="68CCA4E924CC746D4AF1E55A71BC5ED9" pageId="0" pageNumber="301">
The genus 
<taxonomicName id="511A2671793FC4D8746906FA8A8CA1EF" authorityName="Harvey" authorityYear="2004" class="Arachnida" family="Withiidae" genus="Cystowithius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cystowithius" order="Pseudoscorpiones" pageId="0" pageNumber="301" phylum="Arthropoda" rank="genus">
<emphasis id="2BF5B4B832DC33BCF04FD2343BA12FDE" italics="true" pageId="0" pageNumber="301">Cystowithius</emphasis>
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is endemic to Central and South America (
<bibRefCitation id="A59FF356D1FAAF8F3C11D3E6F5F32F62" DOI="https://doi.org/10.1636/S03-48" author="Harvey, MS" journalOrPublisher="The Journal of Arachnology" pageId="0" pageNumber="301" pagination="436 - 456" refId="B15" refString="Harvey, MS, 2004. Remarks on the New World pseudoscorpion genera Parawithius and Victorwithius, with a new genus bearing a remarkable sternal modification (Pseudo-scorpiones, Withiidae). The Journal of Arachnology 32 (3): 436 - 456, DOI: https://doi.org/10.1636/S03-48" title="Remarks on the New World pseudoscorpion genera Parawithius and Victorwithius, with a new genus bearing a remarkable sternal modification (Pseudo-scorpiones, Withiidae)." url="https://doi.org/10.1636/S03-48" volume="32" year="2004">Harvey 2004</bibRefCitation>
; 
<bibRefCitation id="990C36FEAEBE071C73804772D97636A8" DOI="https://doi.org/10.1590/s1984-4689.v38.e21003" author="Garcia, F" journalOrPublisher="Zoologia" pageId="0" pageNumber="301" refId="B13" refString="Garcia, F, Romero-Ortiz, C, 2021. Cystowithius ankeri sp. nov., a new pseudoscorpion from the Central Andes of Colombia (Arachnida: Pseudoscorpiones: Withiidae). Zoologia (Curitiba) 38: e21003. https://doi.org/10.1590/s1984-4689.v38.e21003" title="Cystowithius ankeri sp. nov., a new pseudoscorpion from the Central Andes of Colombia (Arachnida: Pseudoscorpiones: Withiidae). Zoologia (Curitiba) 38: e 21003." url="https://doi.org/10.1590/s1984-4689.v38.e21003" year="2021">
<normalizedToken id="9E8A5A23F3948E49F91C486805B5D2F0" originalValue="García">Garcia</normalizedToken>
and Romero-Ortiz 2021
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), and currently contains five species: 
<taxonomicName id="D65BCDA4A7E1FA7FC71567504E14CC60" lsidName="C. colombicus" pageId="0" pageNumber="301" rank="species" species="colombicus">
<emphasis id="13AE72A46A33227526FD88173353D6E9" italics="true" pageId="0" pageNumber="301">C. colombicus</emphasis>
</taxonomicName>
Harvey, 2004 and 
<taxonomicName id="606880B2DF0FFEB870A0CF3AF2789493" lsidName="C. ankeri" pageId="0" pageNumber="301" rank="species" species="ankeri">
<emphasis id="FED8CEFD60AD2417168B380834564D1D" italics="true" pageId="0" pageNumber="301">C. ankeri</emphasis>
</taxonomicName>
<normalizedToken id="9959D6855C092F1FAA2C7500E547EF4D" originalValue="García">Garcia</normalizedToken>
&amp; Romero-Ortiz, 2021 from Colombia, 
<taxonomicName id="EE4675BD1D63091FB4787CE5F9E4DA73" lsidName="C. ecuadoricus" pageId="0" pageNumber="301" rank="species" species="ecuadoricus">
<emphasis id="D33043265909890FBEF07B574D2A0720" italics="true" pageId="0" pageNumber="301">C. ecuadoricus</emphasis>
</taxonomicName>
(Beier, 1959) and 
<taxonomicName id="0CC35B2B2E2CF28D3D6EE8DBD27038D8" lsidName="C. smithersi" pageId="0" pageNumber="301" rank="species" species="smithersi">
<emphasis id="F76115C8D1573C423A65C8F6526205C1" italics="true" pageId="0" pageNumber="301">C. smithersi</emphasis>
</taxonomicName>
Harvey, 2004 from Ecuador and Peru, and 
<taxonomicName id="BBD70E4033FFE77ED06BDB7707B08DB5" lsidName="C. chamberlini" pageId="0" pageNumber="301" rank="species" species="chamberlini">
<emphasis id="1C503AFE6F45682EC5C2465300A852B9" italics="true" pageId="0" pageNumber="301">C. chamberlini</emphasis>
</taxonomicName>
Harvey, 2004 from Mexico and Guatemala. Males are easily recognised by the presence of sternal invaginations (Figs 
<figureCitation id="9A3F87D7871D57F98DC27877C0C9A852" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Cystowithius florezi sp. nov. (male holotype, ICN-Aps- 077) A carapace B chelicera C left leg I D abdomen, sternites E right chela F left leg IV G left pedipalp H right chelal fingers (arrows show sense-spots) I rallum. Abbreviations: Pedipalp movable finger: b = basal; sb = sub-basal; st = sub-terminal; t = terminal; fixed finger: eb = exterior basal; esb = exterior sub-basal; est = exterior sub-terminal; et = exterior terminal; ib = interior basal; isb = interior sub-basal; ist = interior sub-terminal; it = interior terminal. Cheliceral setae: b = basal seta; es = exterior seta; is = interior seta; ls = laminal seta. Scale bars: 0.5 mm (G); 0.1 mm (A, C-F, H); 0.05 mm (B); 0.0125 mm (I)." figureDoi="10.3897/zookeys.1184.106698.figure5" httpUri="https://binary.pensoft.net/fig/940874" pageId="0" pageNumber="301">5D</figureCitation>
, 
<figureCitation id="8E0403D65BEDA061BFF16738B70D2D01" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Cystowithius ankeri Garcia &amp; Romero-Ortiz, 2021 (male, ICN-Aps- 298) A abdomen sternites B rallum C patch of glandular setae on sternite VIII. Scale bars: 0.1 mm (A); 0.0125 mm (B, C)." figureDoi="10.3897/zookeys.1184.106698.figure6" httpUri="https://binary.pensoft.net/fig/940875" pageId="0" pageNumber="301">6A</figureCitation>
).
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