A new early Late Oligocene (MP 26) continental vertebrate fauna from Saint-Privat-des-Vieux (Alès Basin, Gard, Southern France)
Author
Vianey-Liaud, Monique
Author
Comte, Bernard
Author
Marandat, Bernard
Author
Peigné, Stéphane
Author
Rage, Jean-Claude
Author
Sudre, Jean
text
Geodiversitas
2014
2014-12-26
36
4
565
622
http://dx.doi.org/10.5252/g2014n4a4
journal article
6390
10.5252/g2014n4a4
8bc26961-9853-4113-a63d-4cf691aa7100
1638-9395
4538625
Heterocricetodon gaimersheimensis
Freudenberg, 1941
(
Fig. 12
)
HOLOTYPE
. — Fragment of right lower jaw, bearing m1-3, BSP
1939 XI 4
,
BSPG
, Munich.
TYPE LOCALITY
. — Gaimersheim (South
Germany
),
MP
27-28.
ORIGINAL DIAGNOSIS. — See
Freudenberg (1941: 105)
. Description and variability of teeth of the type-population:
Kristkoiz (1992: 63-72)
.
MEASUREMENTS (in mm). — Eight isolated teeth: 1 M1
(2.54 × 1.66), 3 M2 (2.12 × 1.94, 2.17 × 1.86, 1.94 × 1.79), 1 M3 (1.72 × 1.54),
1 m
1 (2.65 × 1.60),
1 m
2 (2.54 × 1.47),
1 m
3 (2.37 × 1.78).
DESCRIPTION
M1.
Ŋe anterior arm of the protocone and the mesoloph join the labial edge of the tooth, like on several specimens from Gaimersheim.
M2-M3.
Ŋere is a double connecting of the anterior cusps (complete metalophule I and II) and the mesoloph reaches the labial edge of the teeth, as it happens on some specimens from Gaimersheim.
m1.
Ŋe two mesolophids are short, like it is often the case in the
type
population.
m2.
Ŋe posterior arm of the protoconid is short but exceeds the mesolophid length, extending towards the metaconid, as for some specimens from Gaimersheim.
m3.
Ŋe posterior arm of the protoconid is long and joins the labial edge of the tooth, at the base of the metaconid ridge.
DISCUSSION
Four species of relatively large size have been referred to this genus. Among them,
H. stehlini
Schaub, 1925
,
H. schlosseri
(
Schaub, 1925
)
and
H. helbingi
Stehlin & Schaub, 1951
, remain poorly defined.
Hugueney (1980)
, in her discussion about the species of
Heterocricetodon
from Saint-Menoux, has suggested that the variations between the three species
stehlini
,
schlosseri
and
helbingi
could enter a unique species, when more material would be known. Concerning
H. gaimersheimensis
Freudenberg, 1941
,
Engesser (1987)
supposed that “the material from Gaimersheim described by Freudenberg includes two species,
H. stehlini
and
H. helbingi
”. Later, an extensive study has been achieved by
Kristkoiz (1992)
. He established the presence of only one species although with a great variability in the abundant material from Gaimersheim. Ŋerefore,
H. stehlini
and
H. helbingi
could be two morphotypes within
H. gaimersheimensis
and
H. gaimersheimensis
Freudenberg, 1941
would be a junior synonyme of
H. stehlini
Shaub, 1925
. However, the species
H. stehlini
is based on an isolated lower jaw from an undetermined locality from Quercy (“Bach”, Old collections).
FIG. 11.
—
Other cricetids from Saint-Privat-des-Vieux (Gard; Lower Chattian, MP 26):
A
,
Eucricetodon huerzeleri
Vianey-liaud, 1972
, left M1-M2 (reversed), SPV165:
A1
, occlusal view,
A2
, lingual view;
B
,
Cricetidae
indet., fragment of left maxillary with M1, SPV20, occlusal view;
C
, bivariate (Length and Width) diagram of M1 of
Pseudocricetodon incertus
(Schlosser, 1884)
from Saint-Privat-des- Vieux. The open circle represents the M1 SPV20. The plots for the M1 of
P. incertus
are surrounded by their calculated equiprobable ellipse. Scale bar: 1 mm.
FIG. 12.
—
Heterocricetodon gaimersheimensis
Freudenberg,1941
, from Saint-Privat-des-Vieux (Gard;Lower Chattian,MP 26):
A
, right M1 (reversed), SPV170:
A1
, occlusal view,
A2
, lingual view;
B
, left M2, SPV171:
B1
, occlusal view,
B2
, lingual view;
C
, right M3 (re- versed), SPV174:
C1
, occlusal view,
C2
, lingual view;
D
, right m1 (reversed), SPV175:
D1
, occlusal view,
D2
, lingual view;
E
, left m2, SPV178, occlusal view;
F
, left m3, SPV179:
F1
, occlusal view,
F2
, lingual view. Scale bar: 1 mm.
Morphological and dimensional variations of the material of Saint Privat-des-Vieux fall within those of
H. gaimersheimensis
, and are not completely similar with the
type
of
H. stehlini
. Ŋe teeth from Saint- Privat-des-Vieux are thus reported to
H. gaimersheimensis
; because it is the only one species whose variability is known until now. It is the same for a few teeth from Sarèle (Vianey-Liaud 1972) and BoujacA.