Review of the millipede genus Hyleoglomeris Verhoeff, 1910 (Diplopoda, Glomerida, Glomeridae), with descriptions of new species from caves in Southeast Asia Author Golovatch, Sergei I. Institute for Problems of Ecology and Evolution, Russian Academy of Sciences, Leninsky pr. 33, Moscow 119071 (Russia) sgol @ orc. ru sgol@orc.ru Author Geoffroy, Jean-Jacques Muséum national d’Histoire naturelle, Département Écologie et Gestion de la Biodiversité, EA 3778 - USM 306 Réseaux trophiques du sol, 4 avenue du Petit Château, F- 91800 Brunoy (France) geoffroy @ mnhn. fr geoffroy@mnhn.fr Author Mauriès, Jean-Paul Muséum national d’Histoire naturelle, Département Systématique et Évolution, USM 602, case postale 53, 57 rue Cuvier, F- 75231 Paris cedex 05 (France) collmill @ mnhn. fr collmill@mnhn.fr text Zoosystema 2006 28 4 887 915 journal article 10.5281/zenodo.4525415 1638-9387 E114FFC7FFB8FFD32C2E585BCB79FFF4 4525415 3DA8E001-FE3C-418D-9AA1-3CB0BC6F5FD8 Hyleoglomeris maculata n. sp. ( Fig. 6 ) TYPE MATERIAL . — China . Yunnan Province , Menzi County , Lao Shao Dong Cave , 5.I.1989 , leg. P. Beron , holotype ♂ ( NMNHS ), paratypes 4 ♂♂ , 6 ♀♀ , 4 juvs ( NMNHS ) ; 3 ♂♂ , 2 ♀♀ ( MNHN CC154 ) ; 3 ♂♂ , 2 ♀♀ ( ZMUM ). — Same county , Wulichong Sinkhole Cave (No. 3), paratypes 4 ♂♂ , 6 ♀♀ , 4 juvs ( NMNHS ) . ETYMOLOGY. — To emphasize the peculiar maculate colour pattern. DIAGNOSIS. — Differs from congeners by the peculiar colour pattern, combined with a relatively strongly reduced hyposchism and certain details of telopod structure. DESCRIPTION Length 6.0-8.0 ( ♂ ) or 6.5-10 mm ( ♀ ), width 3.0-4.0 ( ♂ ) or 3.3-5.0 mm ( ♀ ). Holotype 7.0 mm long and 3.5 mm wide. Background coloration pallid to yellow, spots on terga brown to dark brown. Head marbled red-brown,antennae dark red-brown, antennomere 6 about 2.3-2.4 times longer than wide; ocelli 6 + 1 to 7 + 1, black, convex;Tömösváry’s organ transverse-oval, only slightly wider than long. Collum, thoracic shield and tergite 11 entirely pale. Body with a distinctive pattern ( Fig. 6A ) of 1 + 1 large, transverse-oval, evidently separated, paramedian spots on terga 3-10 and on pygidium, as well as of 1 + 1 increasingly small, roundish, sublateral spots on terga 3-(8)9. Collum with two transverse striae. Thoracic shield with a narrow hyposchism not reaching the caudal tergal contour; 9 or 10 transverse striae, of which 5 or 6 start above the schism and 4-6 others in differing combinations (never the first and the last though) cross the dorsum. Male pygidium usually slightly sinuate medially at caudal margin. Male leg 17 ( Fig. 6B ) with a medium-sized and regularly rounded outer coxal lobe; telopodite 3- segmented. A D C B FIG. 6. — Hyleoglomeris maculata n. sp. , ♂ paratype from Cave Loa Shao Dong: A , colour pattern; B , leg 17; C , leg 18; D , telopod, front view; E , distal part of telopod, caudal view. Scale bar: A, drawn not to scale; B-E, 0.5 mm. Male leg 18 ( Fig.6C ) with a more or less narrowly ogival syncoxital notch; telopodite 4-segmented. Telopods ( Fig. 6D, E ) with a rather high and subovoid central syncoxital lobe flanked by two setose horns crowned with a foveole-shaped structure with an apical setoid. Prefemur micropapillate laterally. Caudomedial outgrowth of femur relatively narrow at base ( Fig. 6E ). Caudomedial outgrowth of tibia with a micropapillate tubercle at base ( Fig. 6E ). Tarsus weakly rounded apically ( Fig. 6D, E ). REMARKS Based on the coloured head, ocelli and tergal pattern, this species is probably only troglophilic. Its closest relatives seem to be some of the sympatric troglobitic congeners (see below).