The genus Planaeschna McLachlan, 1895 and its subgroupings in Vietnam, with descriptions of three new species (Odonata: Aeshnidae)
Author
Kompier, Tom
Schoutenstraat 69, 2596 SK Den Haag, the Netherlands.
Author
Karube, Haruki
Kanagawa Prefectural Museum of Natural History, Odawara, Japan.
Author
Futahashi, Ryo
National Institute of Advanced Industrial Science and Technology (AIST), Tsukuba, Ibaraki, Japan.
Author
Phan, Quoc Toan
0000-0002-3154-6546
The Center for Entomology & Parasitology Research, College of Medicine & Pharmacy, Duy Tan University, 550000, Da Nang, Vietnam. pqtoan 84 @ gmail. com; https: // orcid. org / 0000 - 0002 - 3154 - 6546
pqtoan84@gmail.com
text
Zootaxa
2021
2021-08-30
5027
1
1
35
journal article
10.11646/zootaxa.5027.1.1
1175-5326
5448029
CCF10500-01A3-476C-A4BE-57161E164EBD
2.
Planaeschna tomokunii
Asahina, 1996
(
Fig. 4
)
Planaeschna tomokunii
:
Asahina (1996)
pp. 70–72,
Figs 4–6
(description of female
holotype
);
Sasamoto
et al.
(2013)
pp. 590–593,
Figs 11
–22 (first description of male, photos of female
holotype
).
Materials examined.
1 ♀
,
Tam Dao NP
,
Vinh Phuc Prov.
,
Vietnam
,
24-IX-2015
, TK leg.;
1 ♂
, same locality and collector,
4-VI-2016
;
1 ♂
, same location and collector,
3-VII-2016
;
2 ♂♂
, same location and collector,
16-VII- 2016
;
1 ♂
1 ♀
, same location and collector,
17-VII-2016
;
1 ♂
, same location and collector,
27-VIII-2016
.
Notes.
Asahina (1996)
described
P. tomokunii
based on a single female. Without detailed explanation, he considered
P. tomokunii
to be closely related to
P. taiwana
. The description of
Planaeschna
species
based on a single female is problematic, because there is little interspecific structural variation in females. For the identification of
Planaeschna
, the combination of the morphology of the male anal appendages and the maculation pattern is effective, but in 1996 there were no
Planaeschna
known from
Vietnam
.
Planaeschna tamdaoensis
with a black and yellow face and plain-faced
P. tomokunii
were the first two species described.
Planaeschna tomokunii
remained enigmatic until
Sasamoto
et al.
(2013)
described the male based on specimens collected at the
type
locality, allowing the species to be correctly identified. So far,
P. tomokunii
may be a true endemic of the Tam Dao mountain range.
Of all
Planaeschna
species
known from
Vietnam
or
China
, none have the combination of slender cerci and dorsal pattern of S2 of male
P. tomokunii
. The dorsal and lateral pattern of S2 is somewhat like that of
P. crux
, but that species has clearly paddle-shaped cerci (see
Fig. 11A–E
). The anal appendages of
P. tomokunii
are similar to those of
P. samurai
and
P. viridis
(compare
Fig. 4F–G
with
Figs 3G–H
and
6B–C
), but differ in detail. The cerci are more parallel-sided in dorsal view compared to
P. samurai
, with a slightly longer epiproct, and in lateral view not waved and curving downward as in
P. viridis
, but gently curving upwards. The dorsal and lateral pattern of S2 distinguishes it clearly from both
P. samurai
and
P. viridis
(compare
Fig. 4D–E
with
Figs 3E–F
and
6D
). Furthermore, the triangular spot on the metepisternum is confluent with the lateral stripe on the mesepimeron in
P. samurai
and
P. viridis
, but separated in
P. tomokunii
(compare
Fig. 4A–C
with
Figs 3B, D
and
6A
).
Ecology.
Planaeschna tomokunii
is found around
1000 m
asl. on the densely forested slopes of Tam Dao, where it apparently breeds in small brooks and seeps. Field observations were done of specimens hawking at dusk over a car track through the forest on the mountainside. In one instance, a male successfully grabbed a female over the track. It occurred together with
P. tamdaoensis
.
Distribution.
Endemic to
Vietnam
(
Vinh Phuc Prov.
)