Systematics of the Hyloxalus bocagei complex (Anura: Dendrobatidae), description of two new cryptic species, and recognition of H. maculosus 2711 Author Páez-Vacas, Mónica I. Author Coloma, Luis A. Author Santos, Juan C. text Zootaxa 2010 2010-12-03 2711 1 75 journal article 1175­5334 Hyloxalus maculosus ECUADOR : Napo : Puerto Misahuallí , Pununo-Jardín Alemán road, 460–462 m , QCAZ 19968–76 , 18878–82 , 20628 , 20630–31 , 20635 , 21186 . Pastaza : Puyo , between Turingia and theatre, 950 m , MCZ-A 91558 ( holotype ) ; 2.6 km W of Puyo on the road, QCAZ 37156–60 ; Canelos , MCZ-A 17938–42 ; 2 km from Rosario Yacu on trail to Union Basi , 99056–57, W of Mera , USNM 164344–45 ; Mera , EPN 1649 , 1656 ; 2 km downstream from headwaters of Río Bobonaza , 655 m , USNM 282067–69 ; headwaters of Río Bobonaza , 685 m , USNM 282070 ; headwaters of Río Arajuno , tributary of Río Napo , USNM 282072 ; Río Pucayacu , USNM 282073–76 ; Río Villano , USNM 282077–79 ; 282683–85; Río Pastaza , USNM 283982 ; 3 km from Shell on Shell-Baños road, QCAZ 37864–65 (tadpoles) , 37868–70; Santa Rosa , 1 km from Shell on Shell-Puyo road, QCAZ 42074–85 , 42086 (tadpoles).