Review of the immature stages of the family Lycidae (Insecta: Coleoptera) Author BOCAK, L. Author MATSUDA, K. text Journal of Natural History 2003 2003-06-30 37 12 1463 1507 http://www.tandfonline.com/doi/abs/10.1080/00222930210125362 journal article 10.1080/00222930210125362 1464-5262 5260079 Calopterini Diagnosis . Calopteron is very similar to Lycostomus (see above). Calopteron differs in entire spiracular plates A1–A8. Caenia differs substantially from both Calopteron and Lycostomus , but only exuviae were available for the study.