Bothynus entellus (LePeletier & Serville) (Coleoptera: Scarabaeidae: Dynastinae) species group: taxonomic revision and description of two new species Author Duarte, Paulo R. M. Author Grossi, Paschoal C. text Zootaxa 2020 2020-03-10 4750 1 101 121 journal article 10.11646/zootaxa.4750.1.5 8c28e10d-e12c-4399-9bd1-bac99bac1fb4 1175-5326 3702869 1A3B7CFF-292B-46AB-82E8-C78BA34618EC Key to males of Bothynus entellus species group Males have pronotal horns, pronounced pronotal excavations, and enlarged anterior claws. 1. Pronotum with 2–3 horns ( Figs. 1A , 3A ), or only with a small ( Fig. 5A ) to moderate ( Fig. 2A ) apical tubercle; if moderate, apical tubercle with rounded apex; mesofemora with setae confine on disc ( Fig. 5F ); tergite VII with 2 bands of weakly marked carinae ( Fig. 7A ); parameres without laterovental contraction at apical half ( Fig. 9A )............................... 2 - Pronotal tubercle strong, horn shaped, acute on apex ( Fig. 4A ); mesofemora completely setose ( Fig. 4B ); tergite VII with 1 band of well-marked carinae ( Fig. 7C ); parameres with lateroventral contraction at apical half in lateral view ( Fig.9D )................................................................................ Bothynus lancifer Dechambre, 1981 2. Pronotum strongly convex posteriorly in lateral view ( Fig. 2B ); pronotum only with an apical tubercle moderate in size; protarsomere IV with elongate ventral apex nearly reaching the apex of the ventro-basal tooth of tarsomere V ( Fig. 2E )......... 3 - Pronotum moderately convex ( Fig. 5B ), or with a horn ( Figs. 1A , 3A ) or tubercle ( Figs. 1C , 3C ) on each side); protarsomere IV with short ventral apex, not reaching the base of the ventro-basal tooth of tarsomere V ( Fig. 5E ).................... 4 3. Metatrochanter with rounded apex ( Fig. 2F ); parameres with subparallel outer margins ( Fig. 8B )............................................................................................ Bothynus stenelus ( Burmeister, 1847 ) - Metatrochanter with acute apex ( Fig. 6B ); parameres with rounded lateral flaps ( Fig. 8F )........................................................................................ Bothynus condacki Duarte & Grossi , new species 4. Pronotum with moderate ( Fig. 1A ) or shallow ( Fig. 5A ) concavity; hypomeron not emarginate in ventral view ( Fig. 5D )... 5 - Pronotal concavity extremely deep ( Fig. 3A ); hypomeron strongly emarginate in ventral view ( Fig. 3E )............................................................................................ Bothynus tricornis Arrow, 1937 5. Stridulatory apparatus with well-marked carinae near to basal margin ( Fig. 7B ); apical half of parameres with subparallel lateral flaps, narrower than basal half ( Fig. 8E ).............................. Bothynus araya Duarte & Grossi , new species - Stridulatory apparatus with weakly marked carinae near to basal margin ( Fig. 7A ); apical half of parameres with rounded lateral flaps, broader than basal half ( Fig. 8A ).............................. Bothynus entellus ( LePeletier & Serville, 1828 )