The history, systematics, and nomenclature of Thalattosuchia (Archosauria: Crocodylomorpha) Author Young, Mark T. School of GeoSciences, Grant Institute, The King’s Buildings, University of Edinburgh, James Hutton Road, Edinburgh, EH 9 3 FE, United Kingdom & LWL-Museum für Naturkunde, Sentruper Strasse 285, 48161 Münster, Germany marktyoung1984@gmail.com Author Wilberg, Eric W. Department of Anatomical Sciences, Stony Brook University, Stony Brook, NY 11794, USA Author Johnson, Michela M. Staatliches Museum für Naturkunde, Rosenstein 1, 70191 Stuttgart, Germany Author Herrera, Yanina Consejo Nacional de Investigaciones Científicas y Técnicas (CONICET), División Paleontología Vertebrados, Museo de La Plata, Facultad de Ciencias Naturales y Museo, Universidad Nacional de La Plata, B 1900 La Plata, Buenos Aires, Argentina. Consejo Nacional de Investigaciones Científicas y Técnicas (CONICET) & Departamento de Diversidade e Ecologia, Faculdade de Biociências, Pontifícia Universidade Católica do Rio Grande do Sul, Av. Ipiranga 6681, Prédio 12, Porto Alegre, 90619 - 900, Brazil & Setor de Paleontologia, Museu de Ciências e Tecnologia, Pontifícia Universidade Católica do Rio Grande do Sul, Av. Ipiranga 6681, Prédio 40, Porto Alegre, 90619 - 900, Brazil Author Brandalise, Marco de Andrade Departamento de Diversidade e Ecologia, Faculdade de Biociências, Pontifícia Universidade Católica do Rio Grande do Sul, Av. Ipiranga 6681, Prédio 12, Porto Alegre, 90619 - 900, Brazil & Setor de Paleontologia, Museu de Ciências e Tecnologia, Pontifícia Universidade Católica do Rio Grande do Sul, Av. Ipiranga 6681, Prédio 40, Porto Alegre, 90619 - 900, Brazil Author Brignon, Arnaud 5, Villa Jeanne-d’Arc, 92340 Bourg-la-Reine, France Author Sachs, Sven Naturkunde-Museum Bielefeld, Abteilung Geowissenschaften, Adenauerplatz 2, 33602 Bielefeld, Germany Author Abel, Pascal Eberhard-Karls-University Tübingen, Fachbereich Geowissenschaften, Sigwartstrasse 10, 72074 Tübingen, Germany Author Foffa, Davide Department of Geosciences, Virginia Tech, 4044 Derring Hall 926 West Campus Drive, Blacksburg 24061, Virginia, USA & School of Geography, Earth and Environmental Sciences, University of Birmingham, Edgbaston, Birmingham B 15 2 TT, United Kingdom Author Fernández, Marta S. Consejo Nacional de Investigaciones Científicas y Técnicas (CONICET), División Paleontología Vertebrados, Museo de La Plata, Facultad de Ciencias Naturales y Museo, Universidad Nacional de La Plata, B 1900 La Plata, Buenos Aires, Argentina. Consejo Nacional de Investigaciones Científicas y Técnicas (CONICET) Author Vignaud, Patrick Laboratoire de Paléontologie, Evolution, Paléoécosystèmes et Paléoprimatologie, CNRS UMR 7262, Department of Geosciences, University of Poitiers, 86073 Poitiers Cedex 9, France Author Cowgill, Thomas School of GeoSciences, Grant Institute, The King’s Buildings, University of Edinburgh, James Hutton Road, Edinburgh, EH 9 3 FE, United Kingdom Author Brusatte, Stephen L. School of GeoSciences, Grant Institute, The King’s Buildings, University of Edinburgh, James Hutton Road, Edinburgh, EH 9 3 FE, United Kingdom text Zoological Journal of the Linnean Society 2024 2024-01-09 200 2 547 617 http://dx.doi.org/10.1093/zoolinnean/zlad165 journal article 10.1093/zoolinnean/zlad165 0024-4082 11241243 1EEF0D52-180B-4D3D-AB95-91AF3091E272 Machimosaurinae Jouve et al ., 2016 (Zoological Code) Machimosaurinae Johnson et al ., 2020a: 118 (PhyloCode) RegNum registration number 759. Type genus: Machimosaurus von Meyer 1837 (emend. von Meyer 1838 ). Etymology ‘Pertaining to Machimosaurus ’. The stem Machimosaur - is based on the genus Machimosaurus . - inae , is a Latin feminine plural suffix for ‘pertaining to’. Under Article 29.2 of the Zoological Code, the suffix - inae denotes a subfamily rank within the family-group. Geological range Middle Jurassic (Bathonian) to Early Cretaceous (Hauterivian or possibly late Barremian?) ( Fanti et al . 2016 , Jouve et al . 2016 , Cortés et al . 2019 , Johnson et al. 2020 a , Young and Sachs 2021). PhyloCode phylogenetic definition Thelargestcladewithin Teleosauroidea containing Machimosaurus hugii ( von Meyer 1837 ) but not Deslongchampsina larteti ( Eudes-Deslongchamps 1866 ) , Macrospondylus bollensis ( Jäger 1828 ) , and Charitomenosuchus leedsi ( Andrews 1909 ) . Definition from Johnson et al . (2022b) . Reference phylogeny Fig. 3 . Zoological Code diagnosis and PhyloCode diagnostic apomorphies Teleosauroid crocodylomorphs with the following unique combination of characters (7): tooth row and occipital condyle aligned in the same plane (shared with the unnamed Chinese teleosaurid and Metriorhynchoidea ) (3.1); ornamentation restricted to the centre of the frontal (shared with the aeolodontin teleosaurids Aeolodon , Bathysuchus and Sericodon ) (22.1); premaxilla–maxilla suture is ‘U’-shaped in dorsal view (shared with the teleosaurid Mystriosaurus ) (132.1); ulna olecranon process greatly expanded (shared with aeolodontin teleosaurids) (768.1); ilium supraacetabular crest shallow and poorly developed (shared with Metriorhynchoidea ) (793.1); ilium postacetabular process expanded into a ‘fan’-shape (795.1); the forelimbs (humerus + ulna) are approximately half the length of the hindlimbs (femur + tibia) at maturity (shared with the hylaeochampsids Pachycheilosuchus and Pietraroiasuchus ) (808.3). Composition Andrianavoay , Neosteneosaurus , Proexochokefalos , and Machimosaurini (composed of Lemmysuchus , Machimosaurus , and Yvridiosuchus ). Comments Authorship: While the nomen Machimosaurinae was first used by Johnson et al . (2020a: 118) , under the Zoological Code Jouve et al. 2016 is the nominal authority. The nominal author of a family-group is the author who first erected a family-group taxon that is valid (in fulfilment of Article 11), and in accordance with the Zoological Code Principle of Coordination applied to family-group names (Article 36.1). However, under the PhyloCode Johnson et al . (2020a) is the nominal authority.