Descriptions of two new species of Bactrocera Macquart (Diptera: Tephritidae: Dacinae: Dacini) from the Philippines with distinct orange medial stripes on the scutum
Author
Korneyev, Severyn V.
0000-0001-8599-7695
California Department of Food and Agriculture, Plant Pest Diagnostics Branch, 3294 Meadowview Road, Sacramento, California, 95832 - 1448, USA. & I. I. Schmalhausen Institute of Zoology, NAS of Ukraine, Bogdan Chmielnicki St. 15, 01030 Kyiv, Ukraine.
Author
Leblanc, Luc
0000-0002-6060-1364
University of Idaho, Department of Entomology, Plant Pathology and Nematology, 875 Perimeter Drive, MS 2329, Moscow, Idaho, 83844 - 2329, USA.
Author
Hauser, Martin
0000-0002-6368-3529
California Department of Food and Agriculture, Plant Pest Diagnostics Branch, 3294 Meadowview Road, Sacramento, California, 95832 - 1448, USA.
Author
General, David E. M.
0000-0001-7735-0023
National Museum of Natural History, National Museum of the Philippines, Malate, Manila, Philippines.
Author
Gaimari, Stephen D.
0000-0002-4240-7154
California Department of Food and Agriculture, Plant Pest Diagnostics Branch, 3294 Meadowview Road, Sacramento, California, 95832 - 1448, USA.
text
Zootaxa
2024
2024-04-03
5432
4
573
599
http://dx.doi.org/10.11646/zootaxa.5432.4.7
journal article
10.11646/zootaxa.5432.4.7
1175-5326
10910781
918057CA-4B08-481B-993A-4A5BA85A6973
Key to
Bactrocera
Macquart
with orange medial stripes on the scutum
1. Scutum without reddish longitudinal stripes,
OR
if present, abdomen without T-shaped dark mark. Wing pattern variable.................................................................................. other species of
Bactrocera
1’. Scutum with one or two submedial orange longitudinal stripes crossing suture
AND
abdomen with a dark T-shaped mark. Wing with well-developed costal band and without crossbands or marks on crossveins r-m and dm-m........................ 2
FIGURE 1.
(a)
B. distincta
(Malloch, 1931)
—wing; (b)
B. usitata
Drew & Hancock, 1994
—scutum, dorsal view; (c)
B. amplexa
(Munro, 1984)
—abdomen, dorsal view; (d-f)
B. ellenriederae
sp. n.
—wing; (e) scutum, dorsal view; (f) abdomen, dorsal view.
FIGURE 2.
Scutum, dorsal view: (a)
B.
(
G
)
continua
(
Bezzi, 1919
)
; (b)
B. lateritaenia
Drew & Hancock, 1994
.
2. Postpronotal lobe connected to lateral postsutural vitta with white area........................................... 3
2’. Postpronotal lobe not connected to lateral postsutural vitta..................................................... 4
3. Lateral margins of abdominal tergite IV yellow. Middle and hind femora entirely yellow.Attractant: no known lure. Distribution:
Philippines
....................................................
Bactrocera
(
Gymnodacus
)
continua
(
Bezzi, 1919
)
3’. Lateral margins of abdominal tergite IV black. Middle and hind femora dark apically. Attractant: cue-lure. Distribution:
Indonesia
. (Illustration available in
Drew & Romig, 2013
)..............................
B
.
(
Ba
.)
pusilla
(Hardy, 1983)
FIGURE 3.
(a–d)
B.
(
G
)
continua
(
Bezzi, 1919
)
—(a) scutum and abdomen, dorsal view; (b) habitus, lateral view; (c) head, frontal view; (d) wing; (e)
B. youngi
sp. n
—
abdomen, dorsal view.
FIGURE 4.
(a–e)
B.
(
Bulladacus
)
mcgregori
(
Bezzi, 1919
)
—(a) scutum, dorsal view; (b) abdomen, dorsal view; (c) head, frontal view; (d) habitus, lateral view; (e) wing; (f)
B. abscondita
(Drew & Hancock, 1981)
—head, frontal view; (g)
B. ellenriederae
sp. n.
—wing.
4. Face without dark spots. Male wing with bulla present on cell cup extension. Attractant: no known lure. Distribution:
Philippines
.....................................
B
. (
Bulladacus
)
mcgregori
(
Bezzi, 1919
)
(or other
Bulladacus
spp.).
4’. Face with two dark spots. Male wing without bulla........................................................... 5
5. Wing infuscation in cell r
4+5
overlapping vein R
4+5
at or nearly at level of R
2+3
vein apex.............................. 6
5’. Wing infuscation in cell r 4+5 overlapping vein R 4+5 significantly distal to level of R 2+3 vein apex....................... 10
FIGURE 5.
Wings: (a)
B. yayeyamana
(Matsumura, 1916)
; (b)
B. tapahensis
Drew & Romig, 2013
; (c)
B. ellenriederae
sp. n.
; (d)
B. dorsalis
(Hendel, 1912)
.
FIGURE 6.
Scutellum: (a)
Zeugodacus tau
(Walker, 1849)
; (b)
B. dorsalis
(Hendel, 1912)
.
6. Scutellum with two pairs of setae......................................................................... 7
6’. Scutellum with one pair of setae......................................................................... 8
7. Thorax with distinct black stripes along entire length of reddish mesonotum. Attractant: no known lure. Distribution:
Japan
(Ryukyu Islands),
Philippines
,
Taiwan
..............................
B.
(
Paratridacus
)
yayeyamana
(Matsumura, 1916)
7’. Thorax dark black with short red stripes.Attractant: no known lure. Distribution:
Bangladesh
,
China
(
Yunnan
),
India
(Andaman Islands),
Indonesia
(
Java
),
Malaysia
(mainland),
Singapore
,
Thailand
....
B.
(
Paratridacus
)
melanius
(
Hardy & Adachi, 1954
)
FIGURE 7.
(a–e)
B. yayeyamana
(Matsumura, 1916)
—(a) scutum & head, dorsal view; (b) abdomen, dorsal view; (c) habitus, lateral view; (d) head, frontal view; (e) wing; (f-j)
B. melanius
(
Hardy & Adachi, 1954
)
(photo previously used in Leblanc
et al.
2021)
—
(f) scutum & head, dorsal view; (g) abdomen dorsal view; (h) habitus, lateral view; (i) head, frontal view; (j) wing.
FIGURE 8.
(a–e)
B. tapahensis
Drew & Romig, 2013
—
(a) scutum & head, dorsal view; (b) abdomen, dorsal view; (c) habitus, lateral view; (d) head, frontal view; (e) wing; (f)
B. dispar
(Hardy, 1982)
—thorax, head & fore legs, lateral view.
8. Fore femur with subapical lateral dark spot. (See also
Drew & Romig, 2013
illustration). Attractant: methyl eugenol. Distribution:
Malaysia
(mainland)........................................
B
.
(
Ba
.)
tapahensis
Drew & Romig, 2013
8’. Fore femur entirely yellow, without a dark spot.............................................................. 9
9. Abdominal tergites IV–V with dark lateral markings. Attractant: methyl eugenol. Distribution:
Indonesia
(
Kalimantan
),
Malaysia
(mainland,
Sabah
), southern
Thailand
.............................
B
(
Ba.
)
unimacula
Drew & Hancock, 1994
9’. Abdominal tergites IV–V without dark lateral markings. Attractant: no known lure. Distribution:
Indonesia
(
Sulawesi
)...................................................................................
B.
(
Ba.
)
dispar
(Hardy, 1982)
FIGURE 9.
(a–e)
B. unimacula
Drew & Hancock, 1994
—
(a) scutum & head, dorsal view; (b) abdomen, dorsal view; (c) habitus, lateral view; (d) head, frontal view; (e) wing; (f–j)
B. dispar
(Hardy, 1982)
—(f) scutum & head, dorsal view; (g) abdomen, dorsal view; (h) habitus, lateral view; (i) head, frontal view; (j) wing.
FIGURE 10.
Thorax, head & fore legs, lateral view: (a)
B. arecae
(
Hardy & Adachi, 1954
)
; (b)
B. bhutaniae
Drew & Romig, 2013
.
10. Fore femur with distinct dark spot........................................................................ 11
10’. Fore femur usually entirely yellow, or with faintly darkened area, but without distinct dark spot....................... 16
FIGURE 11.
(a–d)
B. adamantea
Leblanc &
Doorenweerd, 2018
(photo previously used in Leblanc
et al.
2018)
—
(a) habitus, dorsal view; (b) habitus, lateral view; (c) head, frontal view; (d)—wing; (e)
B. youngi
sp. n.
—scutum, dorsal view.
11. Scutum with yellow apical diamond-shaped medial spot. Attractant: zingerone. Distribution:
Vietnam
...............................................................................
B.
(
Ba.
)
adamantea
Leblanc &
Doorenweerd, 2018
11’. Yellow diamond-shaped medial spot on the scutum absent.................................................... 12
FIGURE 12.
Scutum, dorsal view: (a)
B. rubigina
(Wang & Zhao, 1989)
; (b)
B. dorsalis
(Hendel, 1912)
.
12. Lateral postsutural vitta wedge shaped and ending at or just before intra alar seta.................................. 13
12’. Lateral postsutural vitta parallel sided and overlaps intra alar seta............................................... 14
13. Postpronotal lobe entirely yellow. Mid femur entirely yellow. T-shaped pattern on abdomen well developed. Attractant: no known lure. Distribution:
Cambodia
,
Thailand
,
Vietnam
......................
B.
(
Ba.
)
osbeckiae
Drew & Hancock, 1994
13’. Postpronotal lobe partially darkened along inner margins. Mid femur with dark spot on ventral side. T-shaped pattern on abdomen very narrow and sometimes absent. Attractant: cue-lure. Distribution:
Sri Lanka
..............................................................................................
B
.
(
Ba
.)
selenophora
Tsuruta & White, 2001
FIGURE 13.
(a–e)
B. osbeckiae
Drew & Hancock, 1994
—
(a) scutum & head, dorsal view; (b) abdomen, dorsal view; (c) habitus, lateral view; (d) head, frontal view; (e) wing; (f-j)
B. selenophora
Tsuruta & White, 2001
—(f) scutum & head, dorsal view; (g) abdomen, dorsal view; (h) habitus, lateral view; (i) head, frontal view; (j) wing.
FIGURE 14.
(a–e)
B. carambolae
Drew & Hancock, 1994
—
(a) scutum & head, dorsal view; (b) wing; (c) habitus, lateral view; (d) head, frontal view; (e) abdomen, dorsal view; (f)
B. arecae
(
Hardy & Adachi, 1954
)
—abdomen, dorsal view.
14. Ceromata (spots on tergite V) dark yellow or light brown. Dark pattern on abdominal tergite IV with rectangular projection. Attractant: methyl eugenol. Distribution:
Bangladesh
,
Brunei
,
Cambodia
,
India
(Andaman &
Nicobar Islands
),
Indonesia
(
Java
,
West Nusa Tenggara
),
Malaysia
(mainland,
Sabah
),
Singapore
, southern
Thailand
,
Vietnam
, introduced to northern
Brazil
(
Amapá
), French Guiana,
Guyana
,
Surinam
. Highly invasive pest............
B.
(
Ba.
)
carambolae
Drew & Hancock, 1994
14’. Ceromata (spots on tergite V) dark brown. Dark pattern on abdomen tergite IV without rectangular projection. Attractant: cue-lure or unknown lure.................................................................................. 15
15. Lateral markings on abdominal tergites IV–V diffused and reaching middle of tergite IV and present on tergite V. Attractant: cue-lure. Distribution:
Cambodia
..............................................
B.
(
Ba.
)
kohkongiae
Leblanc, 2015
15’. Lateral markings on abdominal tergites IV–V also diffused but present only on margins of tergite IV, absent on tergite V. Attractant: no known lure. Distribution: Christmas I.,
Malaysia
(mainland),
Singapore
, southern
Thailand
.....................................................................................
B
.
(
Ba
.)
arecae
(
Hardy & Adachi, 1954
)
FIGURE 15.
(a–d)
B. kohkongiae
Leblanc, 2015
(photo previously used in
Leblanc
et al.
2015
a)
—
(a) habitus, dorsal view; (b) habitus, lateral view; (c) head, frontal view; (d)—wing; (e–i)
B. arecae
(
Hardy & Adachi, 1954
)
—
(e) scutum & head, dorsal view; (f) wing; (g) habitus, lateral view; (h) head, frontal view; (i) abdomen, dorsal view.
FIGURE 16.
Wings: (a)
B. occipitalis
(
Bezzi, 1919
)
; (b)
B. dorsalis
(Hendel, 1912)
.
16. Wing with costal band overlapping vein R 2+3 significantly, and as continuous line starting near the point at the end of vein R 1................................................................................................... 17
16’. Wing with costal band confluent with vein R
2+3
not overlapping beyond point at the end of vein R
1
or occasionally faintly overlap............................................................................................. 25
FIGURE 17.
(a) Scutum, dorsal view: (a)
B. dorsalis
(Hendel, 1912)
; (b)
B. youngi
sp. n.
17. Lateral post-sutural vitta parallel sided and reaching intra-alar seta.............................................. 18
17’. Lateral post-sutural vitta wedge shaped and ending or fading out before intra-alar seta.............................. 23
FIGURE 18.
Abdomen, dorsal view: (a)
B. perkinsi
(Drew & Hancock, 1981)
; (b)
B. carambolae
Drew & Hancock, 1994
; (c)
B. occipitalis
(
Bezzi, 1919
)
.
18. T-shaped pattern on abdomen (medial longitudinal black band over tergites) very narrow. Lateral margins of tergites IV–V without significant dark markings........................................................................ 19
18’. T-shaped pattern on abdomen (medial longitudinal black band over tergites) broad. Lateral margins of tergites IV–V with significant dark markings.............................................................................. 20
19. Ceromata (spots on tergite V) dark to black, strongly contrasting with the lighter integument. Attractant: cue-lure. Distribution:
Australia
(
Queensland
, Torres Strait Islands)...............................
B.
(
Ba.
)
perkinsi
(Drew & Hancock, 1981)
19’. Ceromata (spots on tergite V) pale, like the rest of the integument. Attractant: methyl eugenol. Distribution:
Indonesia
(
West Papua
),
Papua New Guinea
,
Australia
(
Queensland
, Torres Strait Is.),
Solomon Islands
. Pest species..................................................................................................
B.
(
Ba.
)
musae
(Tryon, 1927)
FIGURE 19.
(a–e)
B. perkinsi
(Drew & Hancock, 1981)
—(a) scutum & head, dorsal view; (b) abdomen, dorsal view; (c) habitus, lateral view; (d) head, frontal view; (e) wing; (f–i)
B. musae
(Tryon, 1927)
—
(f) habitus, dorsal view; (g) habitus, lateral view; (h) head, frontal view; (i)—wing.
FIGURE 20.
Head, fronto-lateral view: (a)
B. carambolae
Drew & Hancock, 1994
; (b)
B. occipitalis
(
Bezzi, 1919
)
; (c)
B. youngi
sp. n.
20. Face with a pair of oval to circular black spots (length of facial spots
0.25–0.35 mm
; width
0.2–0.25 mm
).............. 21
20’. Face with a pair of elongate oval black spots (length of facial spots
0.26–0.3 mm
; width
0.1–0.12 mm
)................. 22
21. Dark area on scutum in between red stripes without white hairlike microtrichia (for better visibility use dome light). Dark marginal pattern on abdominal tergites III–V usually significantly darkened. Attractant: methyl eugenol. Distribution:
Brunei
,
Indonesia
(
Kalimantan
),
Malaysia
(
Sabah
,
Sarawak
),
Philippines
. Pest species............
B
.
(
Ba
.)
occipitalis
(
Bezzi, 1919
)
21’. Dark area on scutum in between red stripes with white hairlike microtrichia. Dark pattern on abdomen tergite IV with rectangular projection. Attractant: methyl eugenol. Distribution:
Bangladesh
,
Brunei
,
Cambodia
,
India
,
Indonesia
,
Malaysia
,
Singapore
, southern
Thailand
,
Vietnam
; introduced to northern
Brazil
(
Amapá
), French Guiana,
Guyana
,
Surinam
. Highly invasive pest.................................................................
B
. (
Ba
.)
carambolae
Drew & Hancock, 1994
FIGURE 21.
(a–e)
B. occipitalis
(
Bezzi, 1919
)
—
(a) scutum & head, dorsal view; (b) habitus, lateral view; (c) face, fronto-lateral view; (d) abdomen, dorsal view; (e) scutum under high magnification, dorsal view; (f–k)
B. carambolae
Drew & Hancock, 1994
—(f) scutum & head, dorsal view; (g) wing; (h) habitus, lateral view; (i) head, fronto-lateral view; (j) abdomen dorsal view; (k) scutum under high magnification, dorsal view.
FIGURE 22.
(a–e)
B. mediorufula
Drew & Romig, 2013
—
(a) scutum & head, dorsal view; (b) abdomen, dorsal view; (c) habitus, lateral view; (d) head, frontal view; (e) wing; (f–j)
B. lateritaenia
Drew & Hancock, 1994
—(f) scutum & head, dorsal view; (g) abdomen dorsal view; (h) habitus, lateral view; (i) head, frontal view; (j) wing.
22. Lateral postsutural vitta moderately broad, parallel-sided, ending at intra-alar seta. Attractant: methyl eugenol. Distribution:
Indonesia
(
Kalimantan
),
Malaysia
(mainland,
Sarawak
),
Philippines
,
Thailand
....
B.
(
Ba.
)
mediorufula
Drew & Romig, 2013
22’. Lateral postsutural yellow vitta narrow, tapered, sharply narrowing to end at intra-alar seta. Attractant: cue-lure. Distribution:
Brunei
,
Indonesia
(
Sumatra
),
Malaysia
(mainland,
Sabah
),
Vietnam
...........
B.
(
Ba.
)
lateritaenia
Drew & Hancock, 1994
23. Wing pattern in cell r 4+5 covers apical area of wing margin. Facial black spots oval to circular (length of facial spots
0.25 mm
; width
0.18 mm
). Attractant: cue-lure. Distribution:
Bangladesh
,
Bhutan
,
China
(
Guangxi
, Guongdong,
Hainan
),
India
(
Tripura
),
Taiwan
,
Thailand
,
Vietnam
......................................
B
. (
Ba
.)
rubigina
(Wang & Zhao, 1989)
23’. Wing pattern in cell r 4+5 covers most of the wing margin. Face with a pair of elongate oval black spots (length of facial spots
0.26–0.3 mm
; width
0.1–0.12 mm
)...................................................................... 24
FIGURE 23.
(a–e)
B. rubigina
(Wang & Zhao, 1989)
(photo previously used in Leblanc
et al.
2021)—(a) scutum, dorsal view; (b) abdomen, dorsal view; (c) habitus, lateral view; (d) head, frontal view; (e) wing; (f)
B. youngi
sp. n.
—head, frontal view; (g)
B. ellenriederae
sp. n.
—wing.
FIGURE 24.
(a–f)
B. youngi
sp. n.
paratype—(a) scutum & head, dorsal view; (b) habitus, lateral view; (c) head, fronto-lateral view; (d) abdomen dorsal view; (e) male epandrium, lateral view; (f) wing; (g–l)
B. ellenriederae
sp. n.
paratype—(g) scutum & head, dorsal view; (h) habitus, lateral view; (i) head, fronto-lateral view; (j) abdomen dorsal view; (k) male epandrium, lateral view; (l) wing.
24. Subscutellum entirely black. Lateral margins of abdominal tergites III–V with broad black markings and T-shaped pattern shining black with broad medial stripe. Scutum mostly black with two distinct longitudinal red stripes. Surstyli
0.2 mm
long. Attractant: cue-lure. Distribution:
Philippines
(Luzon)........................................
B.
(
Ba.
)
youngi
sp. n.
24’. Red pattern present on subscutellum. Lateral margins of abdominal tergites III–V with narrow dark brown markings and Tshaped pattern dark brown with narrower medial stripe. Scutum black always with reddish-brown areas in addition to two distinct longitudinal red stripes. Surstyli
0.4 mm
long. Attractant: cue-lure. Distribution:
Philippines
(Luzon)............................................................................................
B.
(
Ba.
)
ellenriederae
sp. n.
25. Hind femur yellow with apical portion black. Attractant: cue-lure. Distribution:
Sri Lanka
..............................................................................................
B.
(
Ba.
)
ceylanica
Tsuruta and White, 2001
25’. Hind femur entirely yellow............................................................................. 26
FIGURE 25.
(a–d)
B. ceylanica
Tsuruta and White, 2001
—
(a) habitus, dorsal view; (b) habitus, lateral view; (c) head, frontal view; (d)—wing; (e)
B. cacuminata
Hering, 1941
—
legs & abdomen, lateral view.
FIGURE 26.
Scutum, dorsal view: (a)
B. rubigina
(Wang & Zhao, 1989)
; (b)
B. bhutaniae
Drew & Romig, 2013
.
26. Lateral postsutural vitta wedge shaped, shorter and tapered posteriorly, ending before or barely reaching intra-alar seta.... 27
26’. Lateral postsutural vitta parallel sided, longer, reaching and overlapping intra-alar seta.............................. 28
27. Facial black spots small to medium sized (height of facial spots
0.15 mm
; width
0.18 mm
). Attractant: methyl eugenol. Distribution:
Australia
(
Queensland
,
New South Wales
,
Victoria
, Lord Howe I.)...........
B
(
Ba.
)
cacuminata
Hering, 1941
27’. Facial black spots larger-sized (height of facial spots
0.25 mm
; width
0.18 mm
). Attractant: cue-lure. Distribution:
Bangladesh
,
Bhutan
,
China
(
Guangxi
, Guongdong,
Hainan
),
India
(
Tripura
),
Taiwan
,
Thailand
,
Vietnam
...............................................................................................
B.
(
Ba.
)
rubigina
(Wang & Zhao, 1989)
FIGURE 27.
(a–e)
B. cacuminata
Hering, 1941
—(a) scutum & head, dorsal view; (b) abdomen, dorsal view; (c) habitus, lateral view; (d) head, frontal view; (e) wing; (f–j)
B. rubigina
(Wang & Zhao, 1989)
(photo previously used in Leblanc
et al.
2021)
—
(f) scutum & head, dorsal view; (g) abdomen, dorsal view; (h) habitus, lateral view; (i) head, frontal view; (j) wing.
FIGURE 28.
(a–e)
B. bhutaniae
Drew & Romig, 2013
—
(a) wing; (b) scutum & head, dorsal view; (c) habitus, lateral view; (d) head, fronto-lateral view; (e) abdomen, dorsal view; (f)
B. dorsalis
(Hendel, 1912)
—
abdomen, dorsal view.
28. Ceromata (spots on tergite V) large and reaching medial abdominal stripe.Attractant: cue-lure. Distribution:
Bhutan
,
Cambodia
,
India
(Andaman Islands),
Thailand
,
Vietnam
.................................
B.
(
Ba.
)
bhutaniae
Drew & Romig, 2013
28’. Ceromata (spots on tergite V) smaller and not reaching medial abdominal stripe................................... 29
FIGURE 29.
(a–f)
B. commensurata
Drew & Romig, 2013
—(a) scutum & head, dorsal view; (b) abdomen, dorsal view; (c) habitus, lateral view; (d) head, fronto-lateral view; (e) wing; (f) abdomen, ventral view; (g)
B. dorsalis
(Hendel, 1912)
—
abdomen, ventral view.
29. Abdominal sternum V always pale orange. Attractant: methyl eugenol. Distribution:
Philippines
,
Indonesia
(
Sulawesi
) (
Doorenweerd
et al.
2023
)............................................
B
.
(
Ba
.)
commensurata
Drew & Romig, 2013
29’. Abdominal sternum V from dark orange-brown to brown-black................................................ 30
FIGURE 30.
(a–h)
B. bivittata
Lin & Wang, 2005
—
(a) scutum & head, dorsal view; (b) abdomen, dorsal view; (c) habitus, lateral view; (d) head, fronto-lateral view; (e) scutum under high magnification, dorsal view; (g) wing; (h) abdomen under high magnification, dorsal view; (i–o)
B. dorsalis
(Hendel, 1912)
—(i) scutum & head, dorsal view; (j) wing; (k) habitus, lateral view; (l) head, fronto-lateral view; (m) abdomen, dorsal view; (n) scutum under high magnification, dorsal view; (o) abdomen under high magnification, dorsal view.
30. Dark area on scutum in between of red stripes without white hairlike microtrichia (for better visibility use dome light). Dark marginal pattern on abdominal tergite III always with lateral incision. Attractant: methyl eugenol. Distribution:
Cambodia
,
China
(
Hainan
),
Thailand
(new record)..........................................
B.
(
Ba.
)
bivittata
Lin & Wang, 2005
30’. Dark area on scutum in between of red stripes covered with white microtrichia. Dark marginal pattern on abdominal tergite III not always with lateral incision. [variants with extensive red markings previously known as
B. invadens
Drew, Tsuruta & White, 2005
] Attractant: methyl eugenol. Highly invasive pest. Distribution: Widespread in Africa and South-East Asia, introduced to parts of Oceania.
B. dorsalis
in SE Asia are usually without stripes..........
B.
(
Ba.
)
dorsalis
(Hendel, 1912)