Taxonomic notes on the spider genus Messapus Simon, 1898 (Araneae, Corinnidae), with the description of the new genera Copuetta and Wasaka and the first cladistic analysis of Afrotropical Castianeirinae Author Haddad, Charles R. text Zootaxa 2013 3688 1 1 79 journal article 10.11646/zootaxa.3688.1.1 151481a0-71e1-4402-ab7e-8262caddf539 1175-5326 284201 DB84C679-C2AA-41CC-816E-83E2F6DCA391 Copuetta wagneri sp. nov. Figs 61 , 109 , 151, 152 Etymology: The species is named for German coleopterist Thomas Wagner, in recognition of the many new species of Afrotropical Corinnidae that he has collected by canopy fogging. Diagnosis: This species is closely related to C. maputa sp. nov. Males have an embolus with an oblique base and a narrow elongate tip that is directed at the distal end of the cymbium ( Fig. 109 ). In C. maputa sp. nov. the base is upright and the embolus tip is shorter and directed prolaterally ( Fig. 106 ). Male ( holotype , Budongo, ZMFK). Measurements: CL 3.38, CW 2.62, AL 3.95, AW 2.15, TL 7.35 (6.90– 7.35), FL 0.30, SL 1.51, SW 1.46, AME–AME 0.07, AME–ALE 0.01, ALE–ALE 0.51, PME–PME 0.14, PME– PLE 0.06, PLE–PLE 0.71, PERW 0.97, MOQAW 0.54, MOQPW 0.57, MOQL 0.68. Length of leg segments: I 3.05 + 1.18 + 2.45 + 2.55 + missing = —; II 3.00 + 1.08 + 2.33 + 2.50 + 1.48 = 10.39; III 2.80 + 1.05 + 2.15 + 2.62 + 1.30 = 9.92; IV 3.65 + 1.25 + 2.95 + 3.97 + 1.60 = 13.42. General appearance as in Fig. 61 . Carapace yellow-brown, eye region grey; broad oval pale mottled black marking mediolaterally, separated from lateral fringe by yellow-brown belt, with small narrow oval black marking in front of fovea; striae brown with black mottling, radiating from fovea, directed between coxae; lateral margins with faint narrow black fringe, with large black spots between leg coxae; markings covered with black feathery setae, areas between them covered in white feathery and short straight setae. AME separated by distance slightly less than ½ their diameter; AME separated from ALE by distance less than 1⁄10 AME diameter; clypeus height slightly less than AME diameter; PLE slightly larger than PME; PME separated by distance slightly larger than ½their diameter; PME separated from PLE by distance slightly less than ½ PME diameter; CW: PERW = 2.70:1. Chelicerae yellowbrown, yellow prolaterally at distal end; promargin with two teeth separated by the basal width of proximal tooth, distal tooth much larger; retromargin with two subequal teeth separated by their basal width, distal tooth close to fang base. Endites yellow-brown, cream prolaterally distally; labium brown, cream distally; sternum yellow, brown around borders. Legs yellow-brown, with faint black mottling laterally and black spots at spine bases; markings covered in black feathery setae, with white feathery setae between them; femora I–IV with faint lateral mottling, many small ventral spots and distal ventral band; patellae I–IV with narrow black proximal line and black distal fringe dorsally; tibiae I–III with faint proximal and distal bands at spine pairs, IV with broad median band; metatarsi I–IV with proximal, median and distal bands at paired spines; tarsi without markings. Leg spination: femora: I pl 2 do 3 rl 1, II pl 2 do 3 rl 1, III pl 2 do 3 rl 2, IV pl 2 do 3 rl 1; all femora with scattered erect ventral setae; patellae: I and II with fine proximal and distal do setae, III and IV with proximal seta and distal spine dorsally; tibiae: I pl 1 plv 2 rlv 2, II pl 2 plv 2 rlv 2, III pl 2 do 1 rl 2 plv 2 rlv 1, IV pl 2 do 1 rl 2 plv 2 rlv 1 vt 2; metatarsi: I plv 2 rlv 2, II plv 2 rlv 2, III pl 3 rl 3 plv 2 rlv 2 vt 3, IV pl 3 rl 3 plv 2 rlv 2 vt 3. Palpal spination: femora: pl 1 do 2, with rlv 4 erect setae; patellae: pl 1 do 2; tibiae: pl 1 do 1 plv 1; tarsi: pl 1 plv 3 rlv 1. Abdomen with yellow-brown anterior dorsal scutum extending to ½ abdomen length; dorsum pale grey, covered in yellowbrown and cream feathery and straight setae, with several black chevrons posteriorly; venter pale grey, covered in yellow-brown short straight and feathery setae; epigastric scutum yellow-brown; ventral sclerite absent, single paired row of tiny sclerites laterally; inframamillary sclerite large, subtriangular, dark yellow-brown. Male palp yellow-brown, cymbium brown; tegulum dark orange-brown, with dark red-brown ducts; embolus large, compressed, with two oblique coils and narrow elongate tip directed at the distal end of the cymbium ( Figs 109 , 151, 152 ). FIGURE 153. Distribution of Copuetta lotzi sp. nov. (black circles), C. magna sp. nov. (white circles), C. maputa sp. nov. (black triangles), C. naja sp. nov. (white triangle), C. uzungwa sp. nov. (black star) and C. wagneri sp. nov. (white star) in the Afrotropical Region. Female: Unknown. Type material: Holotype 3: UGANDA : Budongo Forest, 01°45'N , 31°25'E , 1200m a.s.l., leg. T. Wagner, 5– 15.I.1997 (canopy fogging 44, dry season, secondary forest, Rinorea beniensis ) ( ZMFK ). Paratype : Same data, canopy fogging 55, 13 ( ZMFK ). Additional material examined: None. Distribution: Known only from the type locality ( Fig. 153 ). Biology: Collected by canopy fogging in forests.