A review of the genus Pupopsis Gredler, 1898 (Gastropoda: Stylommatophora: Enidae), with the descriptions of eight new species from China Author Wu, Min Author Gao, Linhui text Zootaxa 2010 2725 1 27 journal article 46921 10.5281/zenodo.199982 558b2fa1-879c-44e4-956d-9abd9c9a249a 1175-5326 199982 10. Pupopsis subpupopsis sp. nov. Figs. 1 , 3 H, 5D, 8H–8J, 10; Tables 2 , 3 Type material. Holotype : HBUMM-05676–specimen 1 (fma); Lianghekou, limestone rocks, 1223 m a.s.l., 33°41ʹ47.3ʹ N, 104°29ʹ10.3ʹ E, Zhouqu County, S. Gansu, China , 2006–X–02 , leg. J. Liu & W. Zheng. Paratypes : HBUMM-05650–specimens 1–2 (fms, one broken), limestone rocks, from 1137 m a.s.l., 33°34ʹ21.7ʹ N, 104°39ʹ0 0.3ʹ E to 1151 m a.s.l., 33°34ʹ28.3ʹ N, 104°39ʹ10.7ʹ E, Jiaogong Town, Wudu County, S. Gansu, China , 2006–X–02 , leg. J. Liu & W. Zheng. Type locality. Lianghekou, Wudu County, S. Gansu Material examined. Type material. Distribution. Gansu (only known from the type localitiy). Etymology. The new species is named for its similarity with P . pupopsis . The epithet is a noun in apposition. Diagnosis. Shell with 7.500–7.875 whorls. Height 11.7, diam. maj. 5.1–5.2. Parietal tooth present. Columella with one blunt tooth. Palatal wall with a ridge-like tooth. Epiphallus forming a few loops. Flagellum present. Penial caecum present. Penis distally with two V-shaped pilasters. A-2 near A-3 internally without a ring of papillae. A-3 opening into A-2 without a papilla. Bursa copulatrix duct proximally twisted. Diverticle present. Shell. Cylindrical with conical top; apex not acuminate; dextral; thin-shelled; solid; semitranslucent; glossy. Whorls never spirally grooved; moderately convex. Embryonic shell smooth; more or less polished; with 1.750– 1.750–1.750 whorls. Postnuclear whorls wrinkled. Suture simple. Last whorl ascending immediately behind aperture. Shell 11.7–11.7–11.7 high; 5.1–5.2– 5.2 in diam. maj. Height/diam. maj. ratio 2.26–2.27–2.27. Aperture ovate; slightly oblique; its insertions separated; 3.9–4.2–4.4 high, 3.5–3.6–3.7 wide. Ratio of shell height to aperture height 2.65–2.81–2.97. Palatal margin rounded. Peristome expanded; sharp. Palatal tooth ridge-like. Parietal callus indistinct. Angular tubercle and parietal tooth present. Columellar margin reflexed; with one blunt tooth. Umbilicus a narrow slit. Shell, including upper whorls, uniformly corneous-brown; aperture and teeth whitish. Genitalia. Vas deferens long; of even thickness; entering epiphallus apically. Epiphallus long; cylindrical; forming a few loops. Epiphallic caecum apically blunt. Flagellum present; short; tubular; proximally not expanded; with blunt tip. Penis with terminal entrance of epiphallus; swollen; thin-walled; pilasters fused at epiphallic pore forming velum. With two V-shaped pilasters. V-shaped pilaster with proximal free end approaching at penial retractor insertion. Papilla developed by fused distal pilasters forming V-shaped structure absent. Penial caecum sac-like; situated at boundary between epiphallus and penis. Penial appendix long; branched off from penis at some distance from atrium; divided into sections A-1+A-2, A-3 and A-4+A-5. A-1 long; without verge toward penial chamber. A-1 and A-2 internally with evenly arranged transversal folds. A-1 and A-2 fused. A-2 near A-3 internally without a ring of papillae. A-2 and A-3 not fused. A-3 opening into A-2 without papilla. Boundary between A-4 and A-5 indistinct. A-5 long; straight. Free oviduct moderately long; longer than vagina. Bursa copulatrix duct proximally twisted. Bursa copulatrix small; poorly defined. Diverticle longer than reservoir; slightly expanded. Bursa copulatrix and diverticle distinguishable (diverticle inserted into bursa copulatrix duct at an angle, but bursa copulatrix connected to the duct smoothly); diverticle diverging from bursa duct closer to reservoir than to duct’s proximal end. FIGURE 10. Pupopsis subpupopsis sp. nov. , HBUMM-05676–specimen 1, holotype. A: genitalia, general view; B: distal penis, interior view. Measurement of genitalia. P: 1.1; Ep: 11.6; Fl: 2.1; VD: 13.2; Va: 3; FO: 2.5; BCD: 9.1; BC: 1.0; D: 2.0; A- 1+A-2: 2.5; A-3: 0.9; A-4+A-5: 16.0 ( holotype ). Remarks. This species is conchologically and anatomically closest to P . pupopsis , except that the new species has a diverticle on the bursa copulatrix duct, and the penial caecum is always present.