Five times over: 42 new Angustopila species highlight Southeast Asia's rich biodiversity (Gastropoda, Stylommatophora, Hypselostomatidae) Author Pall-Gergely, Barna https://orcid.org/0000-0002-6167-7221 Centre for Agricultural Research, Plant Protection Institute, Eoetvoes Lorand Research Network, Herman Otto ut 15, H- 1022 Budapest, Hungary pallgergely2@gmail.com Author Hunyadi, Andras Adria setany 10 G 2 / 5., H- 1148 Budapest, Hungary Author Vermeulen, Jaap J. JK Art and Science, Lauwerbes 8, 2318 AT Leiden, Netherlands Author Grego, Jozef Horna Micina 219, SK- 97401 Banska Bystrica, Slovakia Author Sutcharit, Chirasak Animal Systematic Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand Author Reischuetz, Alexander Puechhaimgasse 52, A- 3580 Horn, Austria Author Dumrongrojwattana, Pongrat Department of Biology, Faculty of Science, Burapha University, 169 Longhardbangsaen Road, Muang District, Chonburi, 20131, Thailand Author Botta-Dukat, Zoltan Centre for Ecological Research, Institute of Ecology and Botany, Alkotmany 2 - 4, H- 2600, Vacratot, Hungary Author Oerstan, Aydin 12501 Milestone Manor Lane, Germantown, Maryland, 20876, USA Author Fekete, Judit University of Pannonia, Centre of Natural Science, Research Group of Limnology, Egyetem u. 10, H- 8200 Veszprem, Hungary & Centre for Ecological Research, Institute of Aquatic Ecology, Department of Tisza Research, 18 / c Bem square, H- 4026 Debrecen, Hungary Author Jochum, Adrienne https://orcid.org/0000-0002-6624-6412 Naturhistorisches Museum der Burgergemeinde Bern, CH- 3005 Bern, Switzerland & Institute of Ecology and Evolution, University of Bern, CH- 3012 Bern, Switzerland & Senckenberg Forschungsinstitut und Naturmuseum, 60325 Frankfurt am Main, Germany text ZooKeys 2023 2023-02-13 1147 1 177 http://dx.doi.org/10.3897/zookeys.1147.93824 journal article http://dx.doi.org/10.3897/zookeys.1147.93824 1313-2970-1147-1 9BB9881B0076473D8E53155D37CA1F50 FF2B6B317B505F9EA0E1000BDCD16CE7 Angustopila pustulata Pall-Gergely & Hunyadi sp. nov. Fig. 77 Type material. Holotype : Vietnam • 1 empty shell (H: 0.84 mm, D: 0.93 mm); Hoa Binh Province, Hưng Thi, 37.8 km from Nho Quan towards Hanoi on the Hồ Chi Minh road, left side (locality code: 2020/46); 20°32.74'N , 105°41.89'E ; 15 m a.s.l.; 16 Feb. 2020; A. Hunyadi leg.; HNHM 105312. Paratypes : Vietnam • 1 imaged shell + 58 shells; same data as for holotype; coll. HA • 5 shells; same data as for holotype; coll. JJV. Additional material. Vietnam • 13j/b shells; same data as for holotype; coll. HA. Diagnosis. A medium-sized, conical or (more often) concave-conical Angustopila species with a relatively narrow umbilicus, weak sculpture, comparatively large aperture, a short parietal tooth and a wart-like lower palatal tooth. Description. Shell of normal size for the genus, slightly higher than wide or slightly wider than high; off-white, conical or more often concave-conical; body whorl widest from apertural view; protoconch consists of 1.25 whorls, with weak spiral striation on entire protoconch; teleoconch with fine, dense, irregularly spaced radial growth lines and somewhat stronger, equidistantly and sparsely-arranged spiral lines (ca. 16-19 on body whorl from standard apertural view); sculpture is weaker than in other Angustopila species; whorls 4 or slightly fewer, rounded or slightly shouldered; aperture slightly oblique to shell axis from lateral view; umbilicus relatively narrow; aperture subovoid or reniform with very wide, not distinctly separated sinulus; peristome expanded, not reflected; parietal callus not protruding, not detached from penultimate whorl; parietal tooth straight, strong, relatively low, situated in some distance from peristome edge; lower palatal tooth very small and blunt, blister-like, situated some distance inside the shell from peristome edge. Figure 77. Angustopila pustulata Pall-Gergely & Hunyadi, sp. nov. A paratype B-I holotype (HNHM 105312). Apertural ( A, B ), lateral ( C ), apical ( D ) and ventral ( E ) sides of the shell; sculpture of the protoconch showing protoconch-teleoconch boundary ( F ); aperture ( G ); ventral ( H ) and frontal ( I ) surface of the body whorl. Measurements (in mm). H = 0.84-1.00, D = 0.87-0.93 H/D*100 = 90.3-108.0 ( n = 6), RUD = 21.8-26.9 ( n = 2). Differential diagnosis. Angustopila parallela sp. nov. is larger, has a stronger palatal tooth, stronger spiral striation, and more depressed, more strongly concave shell shape. See also under A. antidomedon sp. nov., A. bathyodon sp. nov., A. gracilis sp. nov. and A. bathyodon sp. nov. Etymology. The specific epithet Angustopila pustulata (blister-bearing in Latin) refers to the low, blister-like palatal tooth. Distribution. Angustopila pustulata sp. nov. is known from the type locality only in Hoa Binh Province of northern Vietnam (Fig. 27 ).