Five times over: 42 new Angustopila species highlight Southeast Asia's rich biodiversity (Gastropoda, Stylommatophora, Hypselostomatidae)
Author
Pall-Gergely, Barna
https://orcid.org/0000-0002-6167-7221
Centre for Agricultural Research, Plant Protection Institute, Eoetvoes Lorand Research Network, Herman Otto ut 15, H- 1022 Budapest, Hungary
pallgergely2@gmail.com
Author
Hunyadi, Andras
Adria setany 10 G 2 / 5., H- 1148 Budapest, Hungary
Author
Vermeulen, Jaap J.
JK Art and Science, Lauwerbes 8, 2318 AT Leiden, Netherlands
Author
Grego, Jozef
Horna Micina 219, SK- 97401 Banska Bystrica, Slovakia
Author
Sutcharit, Chirasak
Animal Systematic Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand
Author
Reischuetz, Alexander
Puechhaimgasse 52, A- 3580 Horn, Austria
Author
Dumrongrojwattana, Pongrat
Department of Biology, Faculty of Science, Burapha University, 169 Longhardbangsaen Road, Muang District, Chonburi, 20131, Thailand
Author
Botta-Dukat, Zoltan
Centre for Ecological Research, Institute of Ecology and Botany, Alkotmany 2 - 4, H- 2600, Vacratot, Hungary
Author
Oerstan, Aydin
12501 Milestone Manor Lane, Germantown, Maryland, 20876, USA
Author
Fekete, Judit
University of Pannonia, Centre of Natural Science, Research Group of Limnology, Egyetem u. 10, H- 8200 Veszprem, Hungary & Centre for Ecological Research, Institute of Aquatic Ecology, Department of Tisza Research, 18 / c Bem square, H- 4026 Debrecen, Hungary
Author
Jochum, Adrienne
https://orcid.org/0000-0002-6624-6412
Naturhistorisches Museum der Burgergemeinde Bern, CH- 3005 Bern, Switzerland & Institute of Ecology and Evolution, University of Bern, CH- 3012 Bern, Switzerland & Senckenberg Forschungsinstitut und Naturmuseum, 60325 Frankfurt am Main, Germany
text
ZooKeys
2023
2023-02-13
1147
1
177
http://dx.doi.org/10.3897/zookeys.1147.93824
journal article
http://dx.doi.org/10.3897/zookeys.1147.93824
1313-2970-1147-1
9BB9881B0076473D8E53155D37CA1F50
FF2B6B317B505F9EA0E1000BDCD16CE7
Angustopila pustulata
Pall-Gergely
& Hunyadi
sp. nov.
Fig. 77
Type material.
Holotype
: Vietnam • 1 empty shell (H: 0.84 mm, D: 0.93 mm);
Hoa
Binh
Province,
Hưng
Thi, 37.8 km from Nho Quan towards Hanoi on the Hồ
Chi
Minh road, left side (locality code: 2020/46);
20°32.74'N
,
105°41.89'E
; 15 m a.s.l.; 16 Feb. 2020; A. Hunyadi leg.; HNHM 105312.
Paratypes
: Vietnam • 1 imaged shell + 58 shells; same data as for holotype; coll. HA • 5 shells; same data as for holotype; coll. JJV.
Additional material.
Vietnam
• 13j/b shells; same data as for holotype; coll. HA.
Diagnosis.
A medium-sized, conical or (more often) concave-conical
Angustopila
species with a relatively narrow umbilicus, weak sculpture, comparatively large aperture, a short parietal tooth and a wart-like lower palatal tooth.
Description.
Shell of normal size for the genus, slightly higher than wide or slightly wider than high; off-white, conical or more often concave-conical; body whorl widest from apertural view; protoconch consists of 1.25 whorls, with weak spiral striation on entire protoconch; teleoconch with fine, dense, irregularly spaced radial growth lines and somewhat stronger, equidistantly and sparsely-arranged spiral lines (ca. 16-19 on body whorl from standard apertural view); sculpture is weaker than in other
Angustopila
species; whorls 4 or slightly fewer, rounded or slightly shouldered; aperture slightly oblique to shell axis from lateral view; umbilicus relatively narrow; aperture subovoid or reniform with very wide, not distinctly separated sinulus; peristome expanded, not reflected; parietal callus not protruding, not detached from penultimate whorl; parietal tooth straight, strong, relatively low, situated in some distance from peristome edge; lower palatal tooth very small and blunt, blister-like, situated some distance inside the shell from peristome edge.
Figure 77.
Angustopila pustulata
Pall-Gergely
& Hunyadi, sp. nov.
A
paratype
B-I
holotype (HNHM 105312). Apertural (
A, B
), lateral (
C
), apical (
D
) and ventral (
E
) sides of the shell; sculpture of the protoconch showing protoconch-teleoconch boundary (
F
); aperture (
G
); ventral (
H
) and frontal (
I
) surface of the body whorl.
Measurements (in mm).
H = 0.84-1.00, D = 0.87-0.93 H/D*100 = 90.3-108.0 (
n
= 6), RUD = 21.8-26.9 (
n
= 2).
Differential diagnosis.
Angustopila parallela
sp. nov. is larger, has a stronger palatal tooth, stronger spiral striation, and more depressed, more strongly concave shell shape. See also under
A. antidomedon
sp. nov.,
A. bathyodon
sp. nov.,
A. gracilis
sp. nov. and
A. bathyodon
sp. nov.
Etymology.
The specific epithet
Angustopila pustulata
(blister-bearing in Latin) refers to the low, blister-like palatal tooth.
Distribution.
Angustopila pustulata
sp. nov. is known from the type locality only in
Hoa
Binh
Province of northern Vietnam (Fig.
27
).