Revision of the rhinoceros beetle genus Oryctophileurus Kolbe with description of a new species, the male of O. varicosus Prell, and notes on biogeography (Scarabaeoidea, Dynastinae, Phileurini) Author Perger, Robert Author Grossi, Paschoal Coelho text ZooKeys 2013 346 1 16 http://dx.doi.org/10.3897/zookeys.346.6114 journal article http://dx.doi.org/10.3897/zookeys.346.6114 1313-2970-346-1 Oryctophileurus armicollis Prell, 1911 Figs 5 A-B ; 6; 7A, C Type material examined. Peru: holotype, male, body length 18.5 mm (ZMHB). Additional material examined. Bolivia: La Paz dep., Nor Yungas prov., Caranavi, I-2003, 2 males with body length 18 mm and 20 mm and 1 female with body length 18 mm (EPGC); Route Coroico-Caranavi, XII-2008, 1 male (EPGC); La Paz dep., Calisaia, V-1925, G.L. Harrington col., 1 male (MZSP); Beni dep., Cosincho, VIII-1925, G.L. Harrington col., 1 female (MZSP). Diagnosis. Color black, strongly shining. Head anteriorly flat, not concave; canthus subquadrate, in some specimens weakly projected forward; horn in males strongly recurved, on about the same level or slightly higher than posterior pronotal protuberance. Mentum with longitudinal furrow flat and wide, as well as posterior margin. Distance between inner teeth of dorsal pronotal protuberance in male wider that width between eyes (Fig. 6A); in females separated by a concavity with occelate punctures; pronotal concavity accounts for about 40-50 % of dorsal pronotal surface in males. Ocellate punctures above posterior-lateral pronotal margin continuous (Fig. 7C). Each elytron with 10 striae; punctures ocellate, sometimes coalescent and elongated, decreasing in size to sides; interstriae smooth, convex. Striae and interstitial punctures arranged in regular lines Mesotibia with 3 to 5 apical teeth; metatibia with 4 to 6 apical teeth. Pygidium in males distinctly more convex than in females, moderately punctate, denser at base and sides. Aedeagus symmetric (Fig. 5A); parameres with apex rounded to subtriangle shaped (Fig. 5B). Figure 6. Dorsal and lateral habitus of Oryctophileurusarmicollis Prell, 1911, A and B holotype male, body length 18.5 mm C and D female, body length 18 mm, scale bar 5 mm. Figure 7. Pronotum of males with body length 20 mm, dorsal, higher areas light, inner teeth of transversal pronotal carina indicated by white arrow: A Oryctophileurus armicollis Prell B Oryctophileurus guerrai ; ocellate punctures at posteriolateral pronotal surface, character state does not differ between sexes C Oryctophileurus armicollis D Oryctophileurus guerrai . Geographical distribution. Oryctophileurus armicollis is known from Peru ( Prell 1911 ; location not specified) and the Andean (Nor Yungas, La Paz dep.) and Subandean (Cosincho, Beni dep.) areas of Bolivia (Fig. 1A). The ecosystem between 800 and 2000 m a.s.l. in this area is considered as South Yungas submontane, subhumid forest ( Josse et al. 2003 ) and receives an annual precipitation between 1500-6000 mm ( Ibisch et al. 2003a ). The locality data suggests that this species is closely associated with the Bolivian Yungas forest and might also occur in the Peruvian Yungas forest.