Dendropaemon Perty, 1830: taxonomy, systematics and phylogeny of the morphologically most derived phanaeine genus (Coleoptera: Scarabaeidae, Scarabaeinae, Phanaeini) Author François Génier Author Patrick Arnaud text Zootaxa 2016 4099 1 journal volume 10.11646/zootaxa.4099.1.1 5d7eab55-66e2-43ed-b764-bac0c6e5b6d6 1175-5326 268274 B73C1BE1-346A-4C89-86B2-51CDACE92877 Dendropaemon (Enicotarsus) Laporte, 1831 Enicotarsus Laporte 1831 , Mag. Zool. 1: pl. 35 (original description) Enicotarsus : Brullé 1837 , In: Hist. Nat . Ins. : 302 (diagnosis) Enicotarsus : Castelnau 1840 , Hist. Nat . Ins. 2: 83 (diagnosis) Enicotarsus : Guérin-Méneville 1844 , Icon. Règ. anim. Cuv. III: 80 (comment) Enicotarsus : Agassiz 1846 , Nom. Zool. : 138 (mention) Enicotarsus : Lacordaire 1856 , Hist. Nat . Ins. III: 102 (mentioned as synonym) Enicotarsus : Burmeister 1861 , Berl. Ent. Zeit. 5: 56 (comment) Enicotarsus : Harold 1869 , Cat. Col. IV: 1020 (mentioned as synonym) Enicotarsus : Gillet 1911 , Col. Cat. 38: 88 (mentioned as synonym) Enicotarsus : Olsoufieff 1924 , Insecta 13: 159 (mentioned as synonym) Enicotarsus : Blackwelder 1944 , U.S. Nat . Mus. Bull. 185: 210 (mentioned as synonym) Enicotarsus : Edmonds 1972 , Univ. Kansas Sc. Bull. 49: 850 (mentioned as synonym) Enicotarsus : Branco 1991 , Ann. Soc. Ent. Fr. (N.S.) 27: 266 (comment) Enicotarsus : Arnaud 2002 , Col. Monde 28: 14 (mentioned as synonym) Type species : Enicotarsus viridipennis Laporte, 1831 ; monotypy. Diagnosis . Size moderate. Entire body except elytra black; elytra with metallic sheen. Body strongly compressed dorsoventrally. Clypeal edge slightly but distinctly emarginate on external side of each clypeal tooth; clypeal teeth more or less triangular. Pronotum only minutely punctate on disc. Elytral base lacking margin. Meso and metatarsi three segmented, first segment elongate, more than three times as long as wide at apex, last segment spiniform, lacing setae apically.