A phylogenetic revision of the true bug genus Heraeus (Hemiptera: Rhyparochromidae: Myodochini), with the description of two new genera and 30 new species Author Dellapé, Pablo M. Author Melo, María C. Author Henry, Thomas J. text Zoological Journal of the Linnean Society 2016 2016-04-17 177 1 29 134 http://dx.doi.org/10.1111/zoj.12362 journal article 10.1111/zoj.12362 0024-4082 10109299 8A961A76-E33C-4171-B401-0B516B62FA6D HERAEUS DOMINICANUS SP. NOV. ( FIGS 3K , 5G , 9I–L , 10 ) Diagnosis Posterior pronotal lobe uniformly brown; meso- and metafemora reddish, pale basally; labium long, extending to metacoxae. The uniformly brown posterior pronotal lobe easily distinguishes H. dominicanus sp. nov. from H. caliginosus . Also, H. dominicanus sp. nov. has a shorter rostrum that extends only to the metacoxae. Description Total length 6.64 ( Fig. 3K ). Head: Strongly convex dorsally, brown, smooth, and shiny, with short recumbent and long erect setae dorsally. Head length 1.55, width 0.99. Postocular length 0.59. Eyes slightly protruding, not surpassing the dorsal margin of head in lateral view. Ocelli placed at level of an imaginary line passing posterior border of eyes. Interocular width 0.50, interocellar width 0.26. Labium light brown with erect setae, extending to meso- or metacoxae (pinned through posterior pronotal lobe). Labial segment lengths: I 0.94, II 1.04, III 0.80, and IV 0.24. Antenna light brown, scapus darker, basiflagellomere reddish distally; with abundant short recumbent and sparse, short, erect setae. Antennal lengths: scape 0.67, pedicel 1.44, basiflagellomere 1.28, and distiflagellomere missing. Thorax: Pronotum dull, pruinose, uniformly brown, punctate, punctures more conspicuous on posterior lobe; setae short and recumbent, anterior pronotal lobe with long erect setae. Collar length 0.11, anterior lobe length 0.78, posterior lobe length 0.56; anterior lobe width 0.99, posterior lobe width 1.55. Pleurae brown. Evaporative area short. Scutellum brown, punctate, pruinose, with long erect setae. Hemelytra pruinose, with short recumbent setae; general colour brown, darkened on posterior half, with a large whitish subapical corial spot. Membrane brown, with an elongate pale apical spot, veins concolorous. Legs: Femora reddish brown, meso- and metafemur paler basally, tibiae pale reddish brown, 52 A REVISION OF HERAEUS ( RHYPAROCHROMIDAE ) Figure 11. Legs: A, Heraeus guttatus ( Dallas, 1852 ) ; B, Heraeus hollyae Baranowski, 2005 ; C, Heraeus steineri sp. nov. ; D, Heraeus triguttatus ( Guérin-Méneville, 1857 ) . tarsi pale brown ( Fig. 5G ). Legs with semi-erect and erect setae; protibia with small spines on inner surface. Abdomen: Brown, shiny, with abundant short recumbent setae. Male genitalia: Pygophore ( Fig. 9I, J ) rounded. Anterior margin of dorsal aperture slightly rounded. Parameres: Figure 9 (K, L). Aedeagus with minute spines, vesical lobes sclerotized; processus gonopori long and slender. Variability observed in a female paratype Distiflagellomeres uniformly light brown, without a pale band. Spines of profemur small and restricted to apical half. Distribution Dominican Republic ( Fig. 10 ). Etymology The specific epithet ‘ dominicanus ’ is an adjective referring to the country where all of the specimens of this species were collected. Type material Holotype : ♂ , DOMINICAN REPUBLIC : S. Frncsco Mts. [ sic ], St. Domingo , W.I. , IX-[20]05, Aug. Busck ( USNM ). Paratype : 1♀ , same data as for holotype ( USNM ) .