† Camelosphecia gen. nov., lost ant-wasp intermediates from the mid-Cretaceous (Hymenoptera, Formicoidea) Author Boudinot, Brendon E. https://orcid.org/0000-0002-4588-0430 Department of Entomology & Nematology, University of California, Davis, One Shields Ave, Davis, CA 95616, USA & Friedrich-Schiller-Universitaet Jena, Institut fuer Zoologie und Evolutionsforschung, 1 Erberstrasse, 07743 Jena, Thueringen, Germany boudinotb@gmail.com Author Perrichot, Vincent https://orcid.org/0000-0002-7973-0430 Univ. Rennes, CNRS, Geosciences - UMR 6118, F- 35000, Rennes, France Author Chaul, Julio C. M. Pos-Graduacao em Ecologia, Departamento de Biologia Geral, Universidade Federal do Vicosa, 36570 - 900, Vicosa, MG, Brazil text ZooKeys 2020 2020-12-18 1005 21 55 http://dx.doi.org/10.3897/zookeys.1005.57629 journal article http://dx.doi.org/10.3897/zookeys.1005.57629 1313-2970-1005-21 B99544631D5746C3AB90751157C9D0B5 610C6AAD64DF5166BE70E5751EC47FD6 Formicoidea Latreille, 1809 Definition. Detailed study of the † Camelomecia clade has redefined the Formicoidea and refined our understanding of the definition and evolutionary patterning of the total and crown Formicidae ( Boudinot et al. 2020a ). Formicoids, we now know, are a clade of Formicapoidina (sister to Apoidea : Johnson et al. 2013 ; Branstetter et al. 2017 ; Peters et al. 2017 ) defined by positive (i.e., non- "absence" character) morphological synapomorphies most of which form an innovation suite for cursorial or surface-based predation, including: (1) prognathy and elongation of the postgenal bridge (Figs 13A , 14B ); (2) enlargement of the dorsal (cranial) mandibular condyle (Fig. 13A ); (3) rotation of the antennal toruli laterad in females (Fig. 13A, B ); (4) elongation of the procoxae (Figs 14A , 15A ); (5) partial to complete enclosure of the proximal protrochanteral articulations within the distal procoxal foramina (Figs 14E , 15A , 16E ); (6) internalization of the proximal meso- and metacoxal articulations within the mesosoma (Figs 14B , 15A , 16 ); (7) petiolation of the first metasomal segment (Figs 14A , 15A , 16A, D, E ); (8) gain of the anteroventral process of the petiolar sternum (Fig. 16A ); (9) buttressing of the metasomal waist through gain of the prora (an anteroventral process of the second metasomal sternum) (Figs 15A , 16A, D ); plus (10) an angled juncture between the first free abscissae of Rs and M in the fore wing (Figs 15A , 16A, D ). The † Camelomecia clade, in contrast to the total clade of the Formicidae , probably lack the metapleural gland and apterous workers altogether, while also being defined by a combination of derived and plesiomorphic features (see, e.g., the key below). Based on direct examination of the unique specimen (holotype) of † Camelomecia janovitzi (BEB at the AMNH, 2017), presence of this gland is uncertain and requires further scrutiny.