New Mycomya species from the Himalayas (Diptera, Mycetophilidae): 2. Subgenera Calomycomya, Cymomya, Neomycomya and Pavomya subg. n. Author Väisänen, Rauno text Zootaxa 2013 3666 3 301 318 journal article 10.11646/zootaxa.3666.3.1 7742af25-2726-4e6d-83b7-3404fa3a3219 1175-5326 216549 5A2C3D51-FB3E-485F-A629-07080EB385D3 Mycomya paguma sp. n. Figs. 6 A–C Material examined. Holotype . 3. NEPAL , 27°56'N , 85°00'E , Mal. tr. 9, 9900', 28.V. 1967 , Can. Nepal Exp. (in CNC). Paratypes (3 only). 28°00'N , 85°00'E , Mal. tr. 7, 9900', 26.V. 1967 , Can. Nepal Exp., 1 3 (in MZH); Gupa Pokali, 27°17'N , 87°30'E , 2900 m—Chouki. 27°12'N , 87°28'E , 2700 m , 16.VI. 1972 , J. Emoto, 1 3 and 1 Ƥ (in KUC). Description . 3. Head. Palp and other mouthparts yellow, clypeus and face brownish, posterior parts of head brown. Antenna brown, scape, pedicel and base of 1st flagellomere yellow. Length of 1st flagellomere about 2.5x, 2nd flagellomere 2x , its width. Thorax. Pronotum yellow, with 3 long setae. Scutum brown, with anterolateral corners yellowish. Anepisternum brownish. Preepisternum brownish. Scutellum brownish, with 4 long setae. Laterotergite brownish. Mediotergite brownish, bare. Wing. Length 3.6–4.1 mm . Wing hyaline. Sc ending in R1 near middle of small cell, Sc1 missing. Apical part of Sc bearing 4–10 small setae. Small cell 1.5–2 times as long as wide. Cu fork slightly proximal to M fork. M ratios: 1.00, 1.38. Cu ratios: 0.66, 1.06. Small setae: M petiole: 0; M1: +; M2: 5; Cu petiole: 0; Cu1: 2–7; Cu2: 2–8. Halter pale yellowish. Legs. Coxae yellow, femora yellow, tibiae and tarsi brownish to brown. Coxa 2 without spur. Leg ratios: bt1:t1 = 0.80–0.85, bt2:t2 = 0.65–0.73, bt3:t3 = 0.58– 0.61. Abdomen. Tergites brown, 3–5 with small yellow anterolateral spots, sternites 1–5 yellow with brownish to brown posterior margins, 6–7 brown. Hypopygium. Figs. 6 A–C, yellow. Sternal lateral appendage short, apically rounded, widest basally ( Fig. 6 B). Sternal submedian filament almost straight, distinctly longer than sternal lateral appendage ( Figs. 6 A–B). Apex of aedeagus rounded in lateral view ( Fig. 6 C). Female. Wing length 4.5 mm . Pronotum yellow. Scutum brown with 2 yellowish longitudinal stripes. Anepisternum light brownish. Preepisternum yellowish, ventral half brownish. Scutellum, laterotergite and mediotergite brownish to brown. Abdominal tergites 1–5 brown with yellowish anterolateral spots, 6–7 brown, sternites lighter. Leg ratios: bt1:t1 = 0.73, bt2:t2 = 0.68, bt3:t3 = 0.58. FIGURE 6. Mycomya paguma sp. n. , male hypopygium. A–B sternal and lateral views, ssf—sternal submedian filament; C apex of aedeagus, lateral view, twice enlarged. Scale 0.2 mm. Discussion . Mycomya paguma differs from the rest of the Himalayan species in having wide, apically rounded sternal lateral appendages ( Fig. 6 B) of the shape of a reversed U (a bit similar to that of the much larger M. panthera ), and the almost straight sternal submedian filaments, which are distinctly longer than the sternal lateral appendages ( Figs. 6 A–B). The male hypopygium of M. paguma resembles that of the Nearctic M. obliqua (Say, 1824) , but differs in the shape of the sternal submedian filaments, which are curved in M. obliqua . Mycomya paguma seems to be closely related to M. gutianshana Wu & Yang, which was described from Zhejiang Province, eastern China , at the altitude of 400–500 m (Wu & Yang 1994). Both species have long and almost straight sternal submedian filaments and the general appearance of the hypopygium is quite similar, but the sternal lateral appendage of M. gutianshana is said to be slender (unfortunately the hypopygium was not figured in a lateral view), while it is wide and rounded in M. paguma . Mycomya gutianshana is smaller (wing length 3.1–3.8 mm ) than M. paguma ( 3.6–4.1 mm ), and its small cell is shorter (1.0–1.5x as long as wide). Although the differences between are M. paguma and M. gutianshana are not great, they are considered to represent two separate species also taking into account that M. paguma is a high altitude species and their known distributions seem to be widely disjunct. Etymology. The species name is a noun in apposition and refers to the Himalayan palm civet (genus Paguma ).