Late Devonian-early Carboniferous bryozoans from Zhankurgan (Greater Karatau, Kazakhstan) - taxonomy and palaeobiogeographical implications Author Tolokonnikova, Zoya A. 0000-0001-6806-4375 Department of oil geology, hydrogeology and geothecnics, Kuban State University, Stavropol’skaya Str., 149, Krasnodar, 350040, Russia. zzalatoi @ yandex. ru; https: // orcid. org / 0000 - 0001 - 6806 - 4375 zzalatoi@yandex.ru Author Fedorov, Petr V. 0000-0002-9888-1381 Department of Sedimentary Geology, Saint Petersburg State University, Universitetskaya nab. 7 - 9, St. Petersburg, 199034, Russia. p. fedorov @ spbu. ru; https: // orcid. org / 0000 - 0002 - 9888 - 1381 p.fedorov@spbu.ru text Zootaxa 2022 2022-10-19 5196 2 252 264 journal article 169973 10.11646/zootaxa.5196.2.6 e618f390-f8c1-4e69-a591-b3b906fef8e6 1175-5326 7224545 45D52870-ADB1-42AA-883D-A1D1D31B5AEF Primorella zhankurganica sp. nov. Figs 4A–F , Table 5 Holotype . SibGIU 12/28 (section with five colony fragments ). Material. SibGIU 12/23.1 (micro-CT). Description. Thin branched colonies. Autozooecial wall 0.02 mm thick in endozone. Diaphragms in the autozooecia absent. Oval autozooecial apertures arranged in diagonal rows; 2 apertures per 1 mm along colony length, 3 diagonally. Autozooecia budding from the central axis at low angles, sharply bending in the exozone and intersecting the colony surface at angles of 80–90°. Abundant aktinotostyles in the walls of the exozone, arranged in regular rows between apertures on the colony surface, displaying distinct core with radially arranged small spines, stellate in cross section. Comparison. Primorella zhankurganica sp. nov. differs from P. variata Tolokonnikova, 2015 from the upper Tournaisian of Kurgan region of Russia (Tolokonnikova 2015) in narrower colonies ( 0.70–0.80 mm vs. 0.67–1.05 mm in P. variata ), abundant aktinotostyles (22–24 around each aperture vs 10–16 in P. variata ) and smaller apertures (0.07–0.08x 0.16–0.20 mm vs. 0.01–0.12x 0.17–0.22 mm in P. variata ). Stratigraphical and geographical ranges. Orgalysay Formation, Mississippian (upper Tournaisian); Karatau, Zhankurgan outcrop, Kazakhstan .