Unraveling a new lineage of Hydrobiidae genera (Caenogastropoda: Truncatelloidea) from the Ponto-Caspian region Author Diana Delicado Author ladimir Pešić Author Peter Glöer text European Journal of Taxonomy 2016 208 1 29 journal article 10.5852/ejt.2016.208 2c5cb62e-9bc7-4fba-a4cb-e5a2fda8c719 2118-9773 269306 urn:lsid:zoobank.org:pub:F807AC79-2615-463B-8272-82523B46210A Genus Intermaria gen. nov. urn:lsid:zoobank.org:act:1CDF6CD1-5D3C-447F-BAAE-7197FBEB 2614 Diagnosis Shell ovate-conic, 3 to 4.5 mm high; large and convex body whorl; rest of whorls small and slightly convex; aperture pyriform, angled on top and often fused to the body whorl. Operculum corneous, yellowish, thin, pliable, ellipsoidal, paucispiral with submarginal nucleus. Two pairs of basal cusps on radular central tooth. Ctenidium occupying nearly the entire length of pallial cavity bearing welldeveloped gill filaments. Osphradium opposite approximate middle of ctenidium. Bursa copulatrix lying against the middle section of the albumen gland; pigmented renal oviduct; one elongate seminal receptacle. Prostate gland bean-shaped, about twice as long as wide; exit of the pallial vas deferens from the posterior-most section of the prostate gland and seminal vesicle entering the prostate gland in its middle section; penis simple, gradually tapering, with its distal end tapered and often with a small distal lobe on the inner edge. Nervous system with black pigmentation typically elongated. Etymology From Latin inter - (between) - maria (seas), referring to the occurrence of the genus in the continental area between the Mediterranean and the Caspian seas. Type species Pseudamnicola zagrosensis Glöer & Pešić, 2009 . Remarks Intermaria species, like those of Pseudamnicola , have an ovate-conic simple shell, one elongated seminal receptacle and a pigmented renal oviduct and nervous system. However, Intermaria differs in having smaller shell dimensions (e.g., when compared with the 5 mm height of P. granjaensis Glöer & Zettler, 2007 see Delicado et al. 2014 ), a more conic-shaped shell, two basal cusps on the central radular tooth (one in Pseudamnicola ), a shorter prostate gland, and a gradually tapering penis with a small distal lobe on the inner edge (penis is triangular with many surface folds and has a blunt end in Pseudamnicola see Delicado et al. 2015 ).