A new genus of millipedes (Diplopoda: Polydesmida: Dalodesmidae) from wet forests in southern Victoria, with brief remarks on the Victorian Polydesmida Author Mesibov, Robert Queen Victoria Museum and Art Gallery, Launceston, Tasmania 7250, Australia mesibov@southcom.com.au text Memoirs of Museum Victoria 2004 2004-12-31 61 1 41 45 https://museumsvictoria.com.au/collections-research/journals/memoirs-of-museum-victoria/volume-61-issue-1-2004/pages-41-45/ journal article 10.24199/j.mmv.2004.61.2 1447-2554 10665547 Victoriombrus gen. nov. Type species. Victoriombrus acanthus sp. nov. , by present designation. Diagnosis . Dalodesmids c. 15 mm long with head + 20 segments, rounded and somewhat swollen paranota and pore formula 5, 7, 9–19. Telopodite of gonopod short, wide, divided by a transverse constriction into a setose, posteromesally excavated basal portion and a bare distal portion bearing a small mesal branch comprising a long, tapering solenomere accompanied by a distally flattened and roughened process, a much larger lateral branch, and between the two a narrow, more or less rod-like process arising posteromesally. Description . Head with moderately deep impression lateral to antennal base; bases separated by about twice a socket diameter; vertigial sulcus faintly indicated; head moderately setose with moderately long setae on clypeus and frons, a few long setae on the vertex. Antenna ( Fig. 1 ) short, slender; antennomere 2 the longest, 6 the widest. Collum about as wide as head and second somite; sparsely setose with moderately long setae; posterior margin straight, anterior margin gently convex, corners rounded. Somite 2 with lateral margin of paranotum well below collum corner, somite 3 margin at about the level of the collum corner. Somites with well-defined waist ( Fig. 1 ); a few setae on somite 2, remaining somites bare and apparently smooth (very small seta-like processes scattered over dorsal surface of metazonite, only visible under high magnification); paranota on most somites just above midlateral ( Fig. 1 ), somewhat swollen, laterally with a very narrow, upwardly concave groove bordered by a slightly thickened margin; from above, anterior margin of paranotum gently curved, lateral margin very slightly convex, posterior margin slightly concave on anterior somites, more so on posterior somites, where the posterior corner projects slightly and bluntly; limbus a comb of fine, straight teeth. Ozopores small, opening dorsolaterally just above and near the posterior end of the paranotal groove on somites 5, 7 and 9–19. Sternites slightly longer than wide, cross impressions well-defined. Anterior legs ( Fig. 1 ) with swollen podomeres, prefemur strongly arched dorsally; posterior legs less swollen; tarsus about as long as next longest podomere, the femur. Sphaerotrichomes on tibia and tarsus only, from leg 3 rearwards, diminishing in number on the more posterior legs and absent from the last leg. Dense ‘brushes’ of setae ventrally on prefemur, femur and postfemur of most legs from 3 rearwards. Leg 2 coxa somewhat extended mesally, the genital opening a simple pit on this extension. Legpairs 4 and 5 not separated at bases, legpairs 6 and 7 well-separated; flexed gonopod telopodites reaching to legpair 6. Spiracles simple in structure with slightly raised rims; on diplosegments the anterior spiracle just above and anterior to the anterior leg, the posterior spiracle just anterior to the midway point between the leg bases. Preanal ring moderately setose; epiproct tapering to blunt, rounded and depressed tip extending well past anal valves; hypoproct paraboloid in outline. Table 1. Native Polydesmida from Victoria.

Taxon	Published localities
Dalodesmidae
Gephyrodesmus cineraceus Jeekel, 1983	Drummer State Forest, 15 km E of Cann River ( Jeekel, 1983 )
Lissodesmus martini ( Carl, 1902 )	Melbourne ( Carl, 1902 ); Ferntree Gully National Park, 18 km NNE of Dandenong ( Jeekel,1983 ); Cockatoo Creek ( Johns, 1964 ).
Paradoxosomatidae
Archicladosoma magnum Jeekel, 1984	Australia ( Jeekel, 1984 ; 1 ); Ferntree Gully National Park, 18 km NNE of Dandenong ( Jeekel, 1984 ; tentative identification)
Australiosoma laminatum Jeekel, 1984	Australia ( Jeekel, 1984 )1
Cladethosoma forceps ( Verhoeff, 1941 )	Gippsland ( Verhoeff, 1941 )
Dicranogonus pix Jeekel, 1982	13 km SE of Buchan; 4 km ESE of Bruthen; Mt Taylor, 11 km NNW of Bairnsdale ( Jeekel, 1982 )
Hoplatessara nigrocingulata Jeekel, 1984	Australia ( Jeekel, 1984 ) 1
H. pugiona Verhoeff, 1941	Whittlesea ( Verhoeff, 1941 )2; Belgrave ( Jeekel, 1984 )
Hoplatria clavigera Verhoeff, 1941	Australia ( Jeekel, 1984 )1; Gippsland ( Verhoeff, 1941 )
Isocladosoma guttatum Jeekel, 1984	Glenaladale National Park, 28 km WNW of Bairnsdale ( Jeekel, 1984 )
I. maculatum Jeekel, 1984	Australia ( Jeekel, 1984 )1
I. pallidulum Jeekel, 1984	Gunyah Gunyah, 32 km SSW of Morwell ( Jeekel, 1984 )
Pogonosternum adrianae Jeekel, 1982	Toongabbie, 18 km NNE of Traralgon ( Jeekel, 1982 )
P. coniferum Jeekel, 1965	Australia ( Jeekel, 1965 , 1982 )1
P. laetificum Jeekel, 1982	Ferntree Gully National Park, 18 km NNE of Dandenong ( Jeekel, 1982 )
P. nigrovirgatum infuscum Jeekel, 1982	Mt Taylor, 11 km NNW of Bairnsdale ( Jeekel, 1982 )
P. nigrovirgatum nigrovirgatum ( Carl, 1902 )	Melbourne ( Carl, 1902 )
Somethus biramus Jeekel, 1984	Drummer State Forest, 15 km E of Cann River; 13 km SE of Buchan; also NSW and SA ( Jeekel, 1984 )

1 Jeekel (1982 , 1984 ) believed that the type specimens of these species are from Victoria because they were collected by Victorian naturalist Charles Barrett and, in the case of Isocladosoma and Pogonosternum spp. , because congeners are known from Victoria . 2 Verhoeff (1941: 14) gives “Whettlerea bei Melbourne”, probably an error for ‘Whittlesea’, c. 40 km north of Melbourne. Gonopod aperture ovoid ( Figs 2 , 4 ), wider than long, about one-half width of somite 7 prozonite, with posterior margin in the form of a flattened ‘V’and posterolateral portions of rim projecting ventrally. Gonocoxa in overall form a truncated cone, tapering distally, with moderately long setae on posteromesal surface; gonocoxae fairly firmly joined along midline but not quite fused into a syncoxite. Cannula prominent. Gonopod telopodites ( Figs 2 , 4 ), short, wide, closely appressed but not joined; partly fused with gonocoxa on anterior side of base; basal portion clearly separated from branched distal portion by a transverse constriction; basal portion excavated posteromesally, sparsely setose, mainly posterolaterally; distal portion bare, divided into a small mesal branch with a long, tapering solenomere joined basally to a distally flattened and roughened process, a much larger lateral branch, and a narrow, more or less rod-like posterior process arising posteromesally. Females somewhat larger than males, with the same colouring; legs smaller than corresponding male legs and not swollen; posterior rim of epigynum projecting slightly, more so in centre; cyphopods not examined. Distribution . Southern Victoria . Etymology . Victoria + Greek ombros (“rainshower”), masculine. Remarks . It is not clear how Victoriombrus is related to other Australian dalodesmid genera. The new genus shares with Tasmanian Bromodesmus ( Mesibov, 2004 ) a long, tapering, acutely pointed solenomere and telopodite elements fringed with teeth, but the gonocoxae in Victoriombrus are more nearly fused into a syncoxite than in the Tasmanian genus, the telopodites are much shorter and wider, and the pore formula is 5, 7, 9–19 instead of 5, 7, 9, 10, 12, 13, 15–19.