Revisiting the spider genus Eutichurus Simon, 1897 (Araneae, Eutichuridae): new species and complementary descriptions Author Bonaldo, Alexandre B. Author Lise, Arno A. Author Ramírez, Martín J. Author Saturnino, Regiane text Zootaxa 2018 2018-02-21 4382 2 321 346 journal article 30692 10.11646/zootaxa.4382.2.6 17f8e584-b4c7-4586-9dbc-c682b3ffcd62 1175-5326 1182079 53C1FBC6-8A60-4C58-A8B9-47311BE186D1 Key to species of Eutichurus (updated from Bonaldo 1994 ) 1 Males (those of E. arnoi , E. manu and E. saylapampa , unknown)................................................ 2 - Females (those of E. abiseo , E. cuzco , E. pallatanga and E. yalen , unknown)..................................... 26 2(1) Retrolateral tibial apophysis represented by a single branch ( Figs 1 , 6, 25; Bonaldo 1994 : figs 19, 29, 38, 48, 56, 64)....... 3 - Retrolateral tibial apophysis bifid or trifid (Figs 14, 17; Bonaldo 1994 : figs 34, 66, 76, 86, 91)........................ 19 3(2) Retrolateral tibial apophysis tuberculate, with a small apical projection ( Bonaldo 1994: figs 56, 57 )....... E. furcifer Kraus - Retrolateral tibial apophysis otherwise ( Figs 1 , 25; Bonaldo 1994 : figs 19, 29, 38, 48, 64)............................ 4 4(3) Retrolateral tibial apophysis short, excavated; median apophysis with a longitudinal median keel ( Bonaldo 1994: figs 43, 45, 48, 49 ).............................................................................................. 5 - Retrolateral tibial apophysis long, not excavated; median apophysis without such keel (Figs 6, 9; Bonaldo 1994 : figs 19, 22, 26, 29, 35, 38, 62)..................................................................................... 6 5(4) Embolus with a prolateral median process ( Bonaldo 1994: figs 43, 44 )........................... E. tropicus (L. Koch) - Embolus without process ( Bonaldo 1994: figs 48, 49 )...................................... E. valderramai Bonaldo 6(4) Embolar base fused to the tegulum ( Fig. 1 ; Bonaldo 1994 : Fig. 35).............................................. 7 - Embolus articulated, connected to the tegulum by a membrane ( Figs 4 , 9 , 12 ; Bonaldo 1994 : figs 19, 22, 30, 26, 38, 62)... 8 7(6) Embolus with lamellar tip ( Bonaldo 1994: figs 35, 36 ).......................................... E. abiseo Bonaldo - Embolus filiform ( Figs 1 , 2)................................................................ E. paredesi n. sp. 8(6) Embolus with a small retrolateral sub-apical process ( Bonaldo 1994 : figs 62, 63; Laborda & Simó 2015 : fig. 1A)................................................................................................ E. ibiuna Bonaldo - Embolus without processes (Figs 5, 10; Bonaldo 1994 : figs 19, 22, 26) or with a prolateral process (Figs 16, 20, 28; Bonaldo 1994 : figs 30, 39)..................................................................................... 9 9(8) Embolus with a prolateral process (Figs 16, 21, 28; Bonaldo 1994 : figs 30, 39).................................... 10 - Embolus without process (Figs 5, 10; Bonaldo 1994 : figs 19, 22, 26)........................................... 15 10(9) Median apophysis strongly sculptured, with several conspicuous sulci ( Bonaldo 1994: figs 29, 31 )........ E. cuzco Bonaldo - Median apophysis not strongly sculptured ( Figs 15 , 20 , 27 , 58 ; Bonaldo 1994 : figs 38, 40).......................... 11 11(10) Embolar prolateral process inserted medially, pars pendula absent (Fig. 28; Bonaldo 1994 : fig. 39).................... 12 - Embolar prolateral process inserted sub-apically, pars pendula present (Figs 21, 59)................................ 13 12(11) Median prong of median apophysis short ( Bonaldo 1994 : figs 38, 40; Ramírez 2014 , figs 147A, C).... E. lizeri Mello-Leitão - Median prong of median apophysis long ( Figs 27 , 29)........................................ E. marquesae Bonaldo 13(11) Embolus wide and short relative to tegulum ( Figs 58 , 59)......................................... E. nancyae n. sp. - Embolus narrow and long relative to tegulum ( Figs 20 , 21, 23, 24)............................................. 14 14(13) Median prong of median apophysis with a pair of apical projections ( Figs 20 , 22)................... E. tequendama n. sp. - Median prong of median apophysis without apical projections ( Figs 23 , 25)......................... E. madre Bonaldo 15(9) Embolus flattened; prolateral margin of median apophysis with transversal ridges ( Bonaldo 1994: figs 26, 28 ).................................................................................................... E. yalen Bonaldo - Embolus conic, median apophysis without transversal ridges (Figs 5, 10; Bonaldo 1994 : figs 19, 22)................... 16 16(15) Retrolateral tibial apophysis expanded distally (Figs 6, 11)................................................... 17 - Retrolateral tibial apophysis tapering toward the apex ( Bonaldo 1994: figs 20, 23 )................................. 18 17(16) Conductor hyaline ( Fig. 4 ).................................................................. E. murgai n. sp. - Conductor partially sclerotized ( Fig. 9 )...................................................... E. zarate Bonaldo 18(16) Retrolateral tibial apophysis gradually tapering from base; embolus inserted basally ( Bonaldo 1994: figs 19, 20 ).................................................................................................... E. ferox Simon - Retrolateral tibial apophysis abruptly tapering from distal third; embolus inserted medially ( Bonaldo 1994: figs 22, 23 )............................................................................................. E. silvae Bonaldo 19(2) Retrolateral tibial apophysis bifid (Figs 14, 17; Bonaldo 1994 : figs 34, 67, 76).................................... 20 - Retrolateral tibial apophysis trifid ( Bonaldo 1994: figs 86, 91 )................................................. 25 20(19) Retrolateral tibial apophysis split medially; median prong of median apophysis present (Fig. 17; Bonaldo 1994 : figs 32, 3 4 ).................................................................................................... 21 - Retrolateral tibial apophysis split basally; median prong of median apophysis absent (Fig. 14; Bonaldo 1994 : figs 67, 72, 76).................................................................................................... 22 21(20) Embolus without prolateral process, tapering distally ( Bonaldo 1994: figs 32, 33 ).................. E. pallatanga Bonaldo - Embolus with prolateral process, distally wide ( Figs 15 , 16)........................................ E. yungas n. sp. 22(20) Embolus conic, without processes ( Bonaldo 1994: figs 75, 76 )..................................... E. luridus Simon - Embolus flattened, with prolateral process (Fig. 13; Bonaldo 1994 : figs 65, 70)................................... 23 23(22) Median apophysis long, with the apex prolaterally oriented; dorsal prong of retrolateral tibial apophysis shorter than the ventral prong ( Figs 12 , 14)....................................................................... E. cumbia n. sp. - Median apophysis short, with the apex oriented proximally; ventral and dorsal prongs of retrolateral tibial apophysis similarly sized ( Bonaldo 1994: figs 67, 72 )........................................................................ 24 24(23) Apex of embolar process acute; apex of embolus apically oriented ( Bonaldo 1994: figs 65–67 )........... E. ravidus Simon - Apex of embolar process rounded; apex of embolus prolaterally oriented ( Bonaldo 1994: figs 70–72 ).. E. itamaraju Bonaldo 25(19) Apex of median apophysis wide; embolar apex bifid ( Bonaldo 1994: figs 89, 91 )........... E. putus O. Pickard-Cambridge - Apex of median apophysis narrow; embolar apex trifid ( Bonaldo 1994: figs 84, 86 )............... E. sigillatus Chickering 26(1) Epigynal posterior plates fused, with a longitudinal median suture (Fig. 18; Bonaldo 1994 : figs 41, 82, 87, 92)........... 27 - Epigynal posterior plates not fused medially (Figs 7, 52; Bonaldo 1994 : figs 21, 46, 59, 68).......................... 31 27(26) Epigynal anterior projections relatively large; median field longer than wide (Fig. 18; Bonaldo 1994 : figs 41, 42)....... 28 - Epigynal anterior projections relatively small; median field wider than long ( Bonaldo 1994: figs 82, 87, 92 )............. 29 28(27) anterior projections covering the anterior half of the posterior plates in ventral view; anterior margin of posterior plates con- verging medially in angle ( Bonaldo 1994 : fig. 41; Ramírez 2014 : fig. 175G)...................... E. lizeri Mello-Leitão - Anterior projections covering less than the anterior half of the posterior plates in ventral view; anterior margin of posterior plates converging medially in a straight line (Fig. 18)............................................. E. yungas n. sp. 29(27) Median field longer than wide, with a posterior median projection; anterior margins of posterior plates W-shaped ( Bonaldo 1994: figs 87, 88 )................................................................... E. sigillatus Chickering - Median field as long as wide, without projection; anterior margins of posterior plates otherwise ( Bonaldo 1994: figs 82, 92 )..................................................................................................... 30 30(29) Anterior projections projected over the median field; anterior margins of posterior plates slightly oblique ( Bonaldo 1994: figs 82, 83 )................................................................................. E. manu Bonaldo - Anterior projections not projected over the median field; anterior margins of posterior plates straight ( Bonaldo 1994: figs 9 2, 93 )......................................................................... E. putus O. Pickard-Cambridge 31(26) Epigynal anterior projections reduced, inconspicuous (Fig. 61)..................................... E. nancyae n. sp. - Epigynal anterior projections conspicuous (Figs 7, 52, 63; Bonaldo 1994 : figs 21, 46, 54).......................... 32 32(31) Anterior projections fused to posterior plates (Figs 63, 65; Laborda & Simó 2015 : fig. 1D)............. E. ibiuna Bonaldo - Anterior projections not fused to posterior plates (Figs 7, 52; Bonaldo 1994 : figs 21, 46, 50)........................ 33 33(32) Each inner margin of anterior projections excavated; posterior plates laterally constricted ( Bonaldo 1994: figs 54, 55 )............................................................................................. E. zarate Bonaldo - Anterior projections and posterior plates otherwise (Figs 7, 52; Bonaldo 1994 : figs 21, 46, 68, 79).................... 34 34(33) Anterior projections larger or slightly smaller than posterior plates (Fig. 7; Bonaldo 1994 : figs 21, 24, 46, 50, 52, 53).... 35 - Anterior projections distinctly smaller than posterior plates ( Fig. 52 ; Bonaldo 1994 : figs 59, 68, 73, 77, 79, 80).......... 41 35(34) Anterior projections triangular, with acute apices (Fig. 7; Bonaldo 1994 : fig. 52).................................. 36 - Anterior projections otherwise ( Bonaldo 1994: figs 21, 24, 46, 50, 53 )........................................... 37 36(35) Posterior margin of anterior projections strongly procurve ( Bonaldo 1994: fig. 52 )................ E. saylapampa Bonaldo - Posterior margin of anterior projections nearly straight (Fig. 7)...................................... E. murgai n. sp. 37(35) Median field subtriangular, bulging between anterior projections ( Bonaldo 1994: fig. 53 )................ E. arnoi Bonaldo - Median field otherwise ( Bonaldo 1994: figs 21, 24, 46, 50 )................................................... 38 38(37) Median field squared, as long as wide; anterior projections truncated ( Bonaldo 1994: fig. 46 )......... E. tropicus (L. Koch) - Median field longer than wide; anterior projections rounded ( Bonaldo 1994: figs 21, 24, 50 )......................... 39 39(38) Apices of anterior projections rugose, strongly sclerotized ( Bonaldo 1994: fig. 50 )............... E. valderramai Bonaldo - Apices of anterior projections smooth ( Bonaldo 1994: figs 21, 24 ).............................................. 40 40(39) Median field surface plain anteriorly, with a longitudinal posterior groove ( Bonaldo 1994: fig. 24 )........ E. silvae Bonaldo - Median field surface concave anteriorly, without groove ( Bonaldo 1994: fig. 21 )........................ E. ferox Simon 41(34) Each posterior plate with a posterior lateral excavation ( Bonaldo 1994: fig. 60 )........................ E. furcifer Kraus - posterior plates without excavations ( Bonaldo 1994: figs 68, 77, 79, 80 )......................................... 42 42(41) Anterior projections projected over the median field ( Bonaldo 1994: figs 68, 73 ).................................. 43 - Anterior projections not projected over the median field ( Bonaldo 1994: figs 77, 79, 80 )............................ 44 43(42) Median field as long as wide ( Bonaldo 1994: fig. 68 )............................................ E. ravidus Simon - Median field longer than wide ( Bonaldo 1994: fig. 73 )....................................... E. itamaraju Bonaldo 44(42) Anterior projections not projected over posterior plates ( Fig. 52 ; Bonaldo 1994 : fig. 80)............. E. marquesae Bonaldo - Anterior projections projected over posterior plates ( Bonaldo 1994: figs 77, 79 )................................... 45 45(44) Anterior projections gently tapering ( Bonaldo 1994: fig. 77 )....................................... E. luridus Simon - Anterior projections truncated ( Bonaldo 1994: fig. 79 ).......................................... E. madre Bonaldo